Andrew Parkinson / Redwings (Turdus iliacus) squabbling over an apple in snow. Derbyshire, UK, February. British Wildlife Photographer of the Year (BWPA) competition 2012, 'Animal Behaviour' category. (Non-ex) Winner of...

Mark Hamblin / 2020VISION / Two Goldfinches (Carduelis carduelis) squabbling over Common teasel (Dipsacus fullonum) seeds in winter, Hope Farm RSPB reserve, Cambridgeshire, England, UK, February. 2020VISION Exhibition.

Mark Hamblin / 2020VISION / Goldfinches (Carduelis carduelis) squabbling over teasel seeds in winter. Hope Farm RSPB reserve, Cambridgeshire, England, UK, February. Crop of 01390677

Colin Varndell / Greenfinches (Carduelis chloris) squabbling in flight, Dorset, UK, February

Laurie Campbell / Goldfinches (Carduelis carduelis) squabbling near seed feeder in garden, Berwickshire, Scotland, April

Laurie Campbell / Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April

Colin Varndell / Male Greenfinches (Carduelis chloris) squabbling in flight. Dorset, UK, March.

Andrew Parkinson / 2020VISION / Marsh harrier (Circus aeruginosus) three sub-adult siblings squabbling in flight in the last rays of evening sunlight, Norfolk, UK, August

Terry Whittaker / 2020VISION / Two Grey herons (Ardea cinerea) squabbling over fish fed to them by visitors,River Tame, Reddish Vale Country Park, Stockport, Greater Manchester, UK, December 2010

Terry Whittaker / 2020VISION / Two Grey herons (Ardea cinerea) on ice, squabbling over fish fed to them by visitors, River Tame, Reddish Vale Country Park, Stockport, Greater Manchester, UK, December 2010, Highly commended, Urban W...

Wild Wonders of Europe / Elander / Bearded vulture (Gypaetus barbatos) adult and juvenile squabbling over food in snow, Cebollar, Torla, Aragon, Spain, November 2008. WWE INDOOR EXHIBITION

Wild Wonders of Europe / Elander / Griffon vultures (Gyps fulvus) squabbling, heads covered in blood from feeding, Andorra, June 2009

Wild Wonders of Europe / Green / Northern gannets (Morus bassanus) on rock squabbling, Saltee Islands, Republic of Ireland, June 2009

Wild Wonders of Europe / Varesvuo / Chaffinch (Fringilla coelebs) two males squabbling at water, Pusztaszer, Hungary, May 2008

Igor Shpilenok / Steller's sea eagle {Haliaeetus pelagicus} adult and juvenile squabbling over Sockeye salmon prey, Kuril Lake, Kamchatka, Far East Russia

Michael Quinton / Trumpeter Swan (Cygnus buccinator) parent and squabbling day-old cygnets, North America

David Pattyn / Dalmatian Pelican (Pelecanus crispus) fighting while a third takes a fish. Lake Kerkini, Greece, March 2012.

Ole Jorgen Liodden / Royal Penguins (Eudyptes schlegeli) squabbling in shallow water, Macquarie Island, Sub-Antarctic Australia. November.

Tui De Roy / Waved Albatross (Phoebastria irrorata) and Blue-footed Booby (Sula nebouxii) squabbling over nesting sites. Galapagos Islands, Ecuador, June.

Mark Hamblin / 2020VISION / Robin (Rubecula eritacus) and coal tit (Parus major) squabbling, Scotland, January

Markus Varesvuo / Four House sparrows (Passer Domesticus) at water, two squabbling, Spain, December

Loic Poidevin / Squabbling European black vultures (Aegypius monachus) as chicks look on in the background. The Pyrenees, Spain, February.

Ernie Janes / Mallard (Anas platyrhynchos) three drakes fighting in water, UK

Ernie Janes / Mallard (Anas platyrhynchos) three drakes squabbling, UK

Ernie Janes / Flock of squabbling Black-headed gulls (Chroicocephalus ridibundus) winter plumage, Norfolk, UK

Bryan and Cherry Alexander / Antarctic / Southern fulmars (Fulmarus glacialoides) squabbling over scraps in the water. Ushuaia, Argentina.

Bryan and Cherry Alexander / Antarctic / Southern fulmars (Fulmarus glacialoides) squabbling over scraps in the water. Ushuaia, Argentina.

Robert Thompson / Razorbills (Alca torda) squabbling on a rock. Great Saltee Island, Kilmore Quay, County Wexford, Republic of Ireland, June.

Terry Whittaker / 2020VISION / Grey herons (Ardea cinerea) squabbling in parkland with cancer charity fun-run participant in background, Regent's Park, London, UK, May 2011.

Danny Green / 2020VISION / Red deer (Cervus elaphus) hind with two Jackdaws (Corvus monedula) squabbling on her head, Richmond Park, London, England, UK, October. Did you know? The crow family is amongst the most intelligent of...

Sergey Gorshkov / Two Kamchatka Brown bears (Ursus arctos beringianus) squabbling beside river, Kamchatka, Far east Russia, July

Sergey Gorshkov / Spotted hyaena (Crocuta crocuta) pair squabbling over carcass, Okavango delta, Botswana, July

Fergus Gill / Northern gannet (Morus bassanus) pair having a dispute with the bird on the neighbouring nest. Troup Head, Scotland, August.

Barrie Britton / African fish eagles (Haliaeetus vocifer) squabbling over carcass of Lesser flamingo (Phoenicopterus minor). Lake Bogoria, Kenya. Picture taken during filming for BBC "Life" TV Series, May 2008

Tim Laman / Two Gentoo penguin chicks (Pygoscelis papua) squabbling, Antarctica, February

Kristel Richard / Feral stallion and his mare squabbling in front of two feral foals, Letea Forest, Danube Delta Biosphere Reserve, Romania, June 2009

Laurie Campbell / Herring Gulls (Larus argentatus) squabbling in flight, Isle of Skye, Hebrides, Scotland, UK, July

Laurie Campbell / Gannets (Morus bassanus), feeding upon fish by catch thrown from fishing boat, Bass Rock, Firth of Forth, Scotland, April

Wild Wonders of Europe / Liodden / Two Black guillemots (Cepphus grylle) squabbling on water, Spitsbergen, Svalbard, March 2009

David Tipling / Steller's sea eagle {Haliaeetus pelagicus} and White-tailed sea eagles {Haliaeetus albicilla} squabbling over fish, Akan, Hokkaido, Japan, February

David Tipling / Gentoo penguin (Pygoscelis papua) incubating adults squabbling in colony, Sea Lion Island, Falklands, November

David Tipling / Two Goldfinches {Carduelis carduelis} squabbling on niger seed feeder in garden, Norfolk, UK

Steve Knell / Northern gannets (Morus bassanus) squabbling at breeding colony, Saltee Islands, County Wexford, Republic of Ireland, May

Andy Rouse / Two Gentoo penguins (Pygoscelis papua) squabbling, Falkland Islands

Andy Rouse / King penguin (Aptenodytes patagonicus) two males squabbling over female at sunset, Falkland Islands

Wild Wonders of Europe / Varesvuo / Hawfinch (Coccothraustes coccothraustes) two males squabbling at water, Pusztaszer, Hungary, May 2008

Wild Wonders of Europe / Varesvuo / Blackcap (Sylvia atricapilla) male squabbling with Common starling {Sturnus vulgaris} at water, Pusztaszer, Hungary, May 2008

Andy Rouse / King penguin (Aptenodytes patagonicus) three males squabbling on beach with storm clouds above, Falkland Islands

Mary McDonald / Giant Petrels {Macronectes giganteus} squabbling over penguin carcass, Falkland Islands

Ian McCarthy / Common starling {Sturnus vulgaris} squabbling over food on roof, Bristol, UK

Suzi Eszterhas / Adelie Penguin (Pygoscelis adeliae) squabbling on iceberg, Paulet Island, Antarctica

Duncan Usher / Northern Fulmar (Fulmarus glacialis) pair squabbling on sea cliff, Northumberland, England

Igor Shpilenok / Steller's sea eagles {Haliaeetus pelagicus} squabbling over Sockeye salmon prey, Kuril Lake, Kamchatka, Far East Russia

Igor Shpilenok / Steller's sea eagles {Haliaeetus pelagicus} squabbling over Sockeye salmon prey, Kuril Lake, Kamchatka, Far East Russia

Steve Knell / Two Gannets (Morus bassanus) squabbling, watched by another in the background. Hermaness National Nature reserve, Shetland Isles, Scotland

Steve Knell / Mixed flock of gulls {Larinae} squabbling at sea over food, Norway, 2006

Anup Shah / Lappet face vulture (Torgos tracheliotus) attacks White backed vultures (Gyps africanus) squabbling over carcass remains, Masai Mara NR, Kenya

Jabruson / Marabou stork (Leptoptilos crumeniferus) squabbling with African fisheagle over flamingo carcass, Lake Nakuru NP, Kenya

Anup Shah / White backed vultures {Gyps africanus} squabbling over carcass, Serengeti NP, Tanzania

Colin Seddon / Great skuas squabbling {Catharacta / Stercorarius Skua} Shetland Isles, Scotland, UK

Colin Seddon / Great skua squabbling {Catharacta / Stercorarius Skua} Shetland Isles, Scotland, UK

GEORGETTE DOUWMA / Grey heron (Ardea cinerea) squabbling. Regent's Park, London, UK.

Paul Hobson / Three Black guillemots squabbling {Cepphus grylle} Scotland

GEORGETTE DOUWMA / Grey herons {Ardea cinerea} squabbling over food Regents Park, London, England

GEORGETTE DOUWMA / Grey herons {Ardea cinerea} squabbling over food, Regents Park, London, England

Pete Oxford / Green winged macaws squabbling at clay lick {Ara chloroptera} Heath river Peru

Ian McCarthy / Giant petrels {Macronectes giganteus} squabbling over food, Marion Island, sub-antarctica

Konrad Wothe / Northern Fulmar (Fulmarus glacialis) pair squabbling, Scotland

Tui De Roy / Sechuran Fox (Lycalopex sechurae) social group members squabbling while foraging at night, Cerro Chaparri, Lambayeque Province, Peru

John Downer / Brown pelicans {Pelecanus occidentalis} squabbling over fish, Stearns Wharf, California, USA

Doug Wechsler / Black Vultures {Coragyps atratus} squabbling and shoving on dead Alligator, Big Cypress NP, Florida, USA