Markus Varesvuo / Osprey (Pandion haliaetus) in flight carrying twig for nest, Vaala, Finland, June

Roger Powell / Short-eared owl (Asio flammeus) hunting, pouncing on prey, Prestwick Carr, Northumberland, UK. December

Rolf Nussbaumer / Red paper wasp (Polistes carolina) adult in flight, Dinero, Lake Corpus Christi, South Texas, USA.

Rolf Nussbaumer / Northern mockingbird (Mimus polyglottos) adult fighting with Golden-fronted Woodpecker (Melanerpes aurifrons) female over Texas Prickly Pear Cactus (Opuntia lindheimeri) Dinero, Lake Corpus Christi, S...

Roland Seitre / Yellow-shouldered Amazon (Amazona barbadensis) in flight. Isla Margarita, Nueva Esparta, Venezuela, 2007.

Fergus Gill / 2020VISION / European Honey Bee (Apis mellifera) with pollen sacs flying towards a Scentless mayweed (Tripleurospermum inodorum) flower in order to feed, Perthshire, Scotland, UK, July. 2020VISION Exhibition. Did...

Andy Rouse / 2020VISION / Red Kite (Milvus milvus) in flight, Wales, UK, November. 2020VISION Exhibition. Did you know? Reintroduced Red kites in England reproduce when they are one year old, but native kites in Wales only sta...

Andrew Parkinson / 2020VISION / A male (left) and female Ptarmigan (Lagopus mutus) in winter plumage taking flight from an ice field as a skier passes by, Cairngorms National Park, Scotland, UK, March 2011. 2020VISION Exhibition.

Danny Green / 2020VISION / Short-eared owl (Asio flammeus) in flight, Worlaby Carr, Lincolnshire, England, UK, December. 2020VISION Exhibition.

Danny Green / 2020VISION / Whooper swans (Cygnus cygnus) in flight, Caerlaverock WWT, Dumfries and Galloway, Scotland, UK. January. 2020VISION Exhibition. 2020VISION Book Plate. Did you know? Fossil records show Whooper swans w...

Peter Cairns / 2020VISION / Northern gannet (Morus bassanus) colony at Hermaness NNR, Unst, Shetland Isles, Scotland, UK, July. 2020VISION Exhibition.

Ben Hall / Whooper swans (Cygnus cygnus) group in flight at dusk, Lancashire, UK December

Ben Hall / Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, 'Habitat' division, British Wildlife Photography Awards (BWPA) competition 2012.

Ben Hall / Whooper swans (Cygnus cygnus) in flight at dusk, Lancashire, UK

Ben Hall / Shelducks (Tadorna tadorna) pair in flight at dusk, Lancashire, UK

Fabrice Cahez / Eurasian griffon vulture (Gyps fulvus) Gorges du Tarn, France, January.

Fabrice Cahez / Eurasian griffon vulture (Gyps fulvus) three in flight, Gorges de la Jonte, France, January.

Fabrice Cahez / Eurasian griffon vulture (Gyps fulvus) in flight, Gorges de la Jonte, France, January.

Fabrice Cahez / Eurasian griffon vulture (Gyps fulvus) in flight, diving downwards, Gorges du Tarn, France, January.

Fabrice Cahez / Eurasian griffon vulture (Gyps fulvus) in flight above clouds, Gorges du Tarn, France, January.

Fabrice Cahez / Eurasian griffon vulture (Gyps fulvus) in flight above clouds, Gorges du Tarn, France, January.

Ross Hoddinott / 2020VISION / Southern hawker dragonfly (Aeshna cyanea) in flight, Arne RSPB reserve, Dorset, England, UK, August. 2020VISION Book Plate.

Ross Hoddinott / 2020VISION / Male Common Darter dragonfly (Sympetrum striolatum) in flight, Arne RSPB reserve, Dorset, England, UK, August. 2020VISION Book Plate.

Ernie Janes / Greylag geese (Anser anser) in flight with paraglider behind, Cley bank, Norfolk, UK

David Kjaer / Flock of Common Starling (Sturnus vulgaris) gathering before landing at winter roost, Salisbury Plain, Wiltshire, UK, January

Marie Read / Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February

Roy Mangersnes / Glaucious gull (Larus hyperboreus) flying against a blue glacier front, Spitsbergen, Svalbard, Norway, September

Roy Mangersnes / Black-legged kittiwakes (Rissa tridactyla) in flight during evening snowfall, Northern Nordaustlandet, Svalbard, Norway, August

Chris Gomersall / 2020VISION / Buff-tailed bumble bee (Bombus terrestris) worker alighting on a Scorpionweed (Phacelia tancetifolia) flower in a conservation margin in order to feed, Hope Farm RSPB reserve, Cambridgeshire, England,...

Chris Gomersall / 2020VISION / A flock of Pink-footed geese (Anser brachyrhynchus) taking flight from a sugar beet field in Norfolk, England, UK, November

Roy Mangersnes / Northern goshawk (Accipiter gentilis) chasing a Hooded crow (Corvus cornix) Southern Norway, December

Roy Mangersnes / Northern Goshawk (Accipiter gentilis) chasing a Hooded crow (Corvus cornix) Southern Norway, December

Roy Mangersnes / Atlantic puffin (Fratercula arctica) in flight against snow, Finnmark, Northern Norway, April

Roy Mangersnes / Herring gull (Larus argentatus) in flight with others on water, Norway, July

Roy Mangersnes / Two Barnacle geese (Branta leucopsis) in flight, Spitsebergen, Svalbard, Norway, August

Roy Mangersnes / Juvenile Black-legged kittiwake (Rissa tridactyla) flying over Raudfjorden, North-western tip of Spitsbergen, Svalbard, Norway, September

David Tipling / 2020VISION / Small flock of Greylag geese (Anser anser) flying over snow covered fields, Ken-Dee Marshes RSPB reserve, Dumfries and Galloway, Scotland, UK, December

Chris Gomersall / 2020VISION / Monochrome image of a Northern gannet (Morus bassanus) adult in flight, seen from the pelagic trawler 'Charisma', Shetland Isles, Scotland, UK, October

Chris Gomersall / 2020VISION / Adult Kittiwake (Rissa tridactyla) in winter plumage in flight alongside the pelagic trawler 'Charisma', Shetland Isles, Scotland, UK, October

Chris Gomersall / 2020VISION / Fulmar (Fulmarus glacialis) in flight over rough sea alongside the pelagic trawler 'Charisma' in evening light, Shetland Isles, Scotland, UK, October. 2020VISION Book Plate.

Chris Gomersall / 2020VISION / Northern gannet (Sula bassana) being mobbed by immature Great black-backed gull (Larus marinus), Shetland Islands, Scotland, UK, October

Mike Read / Cranes (Grus grus), imprinted from hatching to fly with a microlight. Hampshire County Show near Brockenhurst, Hampshire, England, July 2008.

Chris Gomersall / 2020VISION / Mixed flock of Northern gannets (Morus bassanus) and gulls (Larus) feeding in the wake of the pelagic trawler 'Charisma' in evening light, Shetland Islands, Scotland, UK, October 2011

Nick Turner / Boy kite flying on Selsley Common (unimproved grassland) Stroud, Gloucestershire, UK, August 2011. Model released

Alex Mustard / Mirrorwing flying fish (Hirundichthys speculiger) mirrored and swimming beneath the surface at night, between Bahamas and Gulf Stream, West Atlantic Ocean, Florida, USA.

Laurie Campbell / Goldfinches (Carduelis carduelis) squabbling near seed feeder in garden, Berwickshire, Scotland, April

Konrad Wothe / Flock of Snow Geese (Chen caerulescens atlanticus / Chen caerulescens) in flight to their winter roosting site, crossing a full moon, Bosque del Apache, New Mexico, USA, November

Konrad Wothe / Three Snow Geese (Chen caerulescens atlanticus / Chen caerulescens) in flight, Bosque del Apache, New Mexico, USA, November

Laurie Campbell / Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April

Laurie Campbell / Black-headed gulls (Larus ridibundus) robbing Puffin (Fratercula arctica) of catch, Farne Islands, Northumberland, England, June

Laurie Campbell / Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March

Terry Whittaker / 2020VISION / Flock of Pale-bellied brent geese (Branta bernicla) flying over town of Burnham-on-Crouch, Wallasea Wild Coast Project, RSPB Greater Thames Futurescapes Project, Wallasea Island, Essex, England, UK, N...

Terry Whittaker / 2020VISION / Short-eared owl (Asio flammeus) hunting over farmland, with Burnham-on-Crouch in background, Wallasea Wild Coast Project, RSPB Greater Thames Futurescapes Project, Wallasea Island, Essex, England, UK,...

Terry Whittaker / 2020VISION / Short-eared owl (Asio flammeus) hunting over farmland, with Burnham-on-Crouch in background, Wallasea Wild Coast Project, RSPB Greater Thames Futurescapes Project, Wallasea Island, Essex, England, UK,...

Terry Whittaker / 2020VISION / Lesser black-backed gulls (Larus fuscus), Herring gulls (Larus argentatus), Black-headed gulls (Larus ridibundus) and Starlings (Sturnus vulgaris) feeding on landfill site, Pitsea, Essex, England, UK,...

Terry Whittaker / 2020VISION / Mixed flock of Gulls (Larus sp.) flying over a landfill site, Pitsea, Essex, England, UK

Terry Whittaker / 2020VISION / Mixed flock of Gulls (Larus sp.) flying over landfill site, Pitsea, Essex, UK, November 2011

Terry Whittaker / 2020VISION / Lesser black-backed gulls (Larus fuscus), Herring gulls (Larus argentatus), Black-headed gulls (Larus ridibundus) and Starlings (Sturnus vulgaris) feeding on landfill site, Pitsea, Essex, England, UK,...

Terry Whittaker / 2020VISION / Lesser black-backed gulls (Larus fuscus), Herring gulls (Larus argentatus), Black-headed gulls (Larus ridibundus) and Starlings (Sturnus vulgaris) feeding on landfill site, Pitsea, Essex, England, UK,...

Bertie Gregory / 2020VISION / Adult peregrine falcon (Falco peregrinus) flying next to a tower block, London, England, UK, June

Andrew Parkinson / 2020VISION / An adult male Mallard (Anas platyrhynchos) comes in to land, backlit by evening sunlight, Derbyshire, England, UK, March. Did you know? In addition to the colour differences between male and female du...

Peter Cairns / 2020VISION / Arctic Tern (Sterna paradisaea) against white sky, Farne Islands, Northumberland, England, UK, July

Bertie Gregory / 2020VISION / Adult Peregrine falcon (Falco peregrinus) flying near a tower block, London, England, UK, May

Bertie Gregory / 2020VISION / Juvenile Peregrine falcon (Falco peregrinus) flying in front of a Union Jack flag on top of the Houses of Parliament, London, England, UK, July. 2020VISION Exhibition. 2020VISION Book Plate. Did you k...

Bertie Gregory / 2020VISION / Juvenile Peregrine falcons (Falco peregrinus) chasing one another in front of a Union Jack flag on top of the Houses of Parliament, London, England, UK, July

Sergey Gorshkov / Willow grouse / ptarmigan (Lagopus lagopus) pair in flight, Agapa River, Taimyr Peninsula, Siberia, Russia

Brent Stephenson / Flock of Cape petrels (Daption capense capense) in flight against the sea. Against the sun. South Shetland Islands, Antarctica, January.

Brent Stephenson / Wilson's Storm-petrel (Oceanites oceanicus) hovering whilst feeding in calm conditions. Cuverville Island, Antarctic Peninsula, Antarctica, December.

Brent Stephenson / Flock of Cape Petrels (Daption capense capense) in flight, heading away against the sky. South Shetland Islands, Antarctica, December.

Brent Stephenson / Light-mantled Sooty Albatross (Phoebetria palpebrata) close up in flight at eye level, portrait. Drake Passage, South Atlantic, December.

Brent Stephenson / New Zealand Albatross (Diomedea antipodensis) in flight, showing upperwing, wingtip touching the sea surface. Off Kaikoura, Canterbury, New Zealand, October.

Brent Stephenson / Male Australasian Gannet (Morus serrator) flying with seaweed for use as nesting material. Cape Kidnappers, Hawkes Bay, New Zealand, September.

Todd Mintz / Great Horned owl, (Bubo virginianus) in flight, Regina, Saskatchewan, Canada, March

Markus Varesvuo / Barn swallow feeding chick on the wing (Hirundo rustica) Sipoo Finland July

Markus Varesvuo / Spotted Redshank (Tringa erythropus) in flight, Utsjoki Finland June

Markus Varesvuo / Magpie (Pica pica) in flight, Sweden October

Markus Varesvuo / Dunlin (Calidris alpina) flock in flight, Finland July

Nick Upton / 2020VISION / Honey bee (Apis mellifera) hovering near Snowberry flowers (Symphoricarpos sp.) in Parade gardens park, with city buildings in the background, Bath, England, UK, June

Pete Oxford / Sparkling violetear hummingbird (Colibri coruscans) at base of Chimborazo Volcano (Highest mountain in Ecuador), Andes, Ecuador, South America

Steve Gettle / Violet-crowned Woodnymph (Thalurania colombica) hummingbird male flying, Costa Rica

Nick Upton / 2020VISION / Flock of Wigeon (Anas penelope) and a few Common Teal (Anas crecca) flying overhead in winter, Greylake RSPB reserve, Somerset Levels, UK, December

Nick Upton / 2020VISION / Lapwings (Vanellus vanellus), Golden plover (Pluvialis apricaria), Wigeon (Anas penelope) and Common Teal (Anas crecca) flying over flooded marshes in winter, with a rainbow in the background, Greylak...

Edwin Giesbers / Two Scarlet macaws (Ara macao) flying, captive

Chadden Hunter / Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009

Thomas Rabeil / Flock of Marabou Stork (Leptoptilos crumeniferus) taking flight at seasonal waterhole, seen from above. Sahelo-Sudanese Biome, W National Park (UNESCO, IUCN & RAMSAR), Niger. Aerial census, May 2011.

John Abbott / Eastern Green june beetle (Cotinis nitida) in flight, Texas, USA, October

John Abbott / Snakefly (Agulla sp) in flight, Wild Basin Preserve, Texas, USA, April

Loic Poidevin / Flock of Oystercatchers (Haematopus ostralegus) flying in formation. Breton Marsh, French Atlantic Coast, August.

Loic Poidevin / Flock of Common Crane (Grus grus) in flight silhouetted against the sun. France, November.

Loic Poidevin / Adult Common Tern (Sterna hirundo) hovering over its chicks, one of which has taken a small fish from the adult. Noirmoitier Island, French Atlantic Coast, July.

Juan Carlos Munoz / Jersey tiger moth (Euplagia quadripunctaria) large numbers resting on the trunk of Sweetgum trees (Liquidambar orientalis) some with wings open in flight, in the Petaloudes Valley, Rhodes island, Gree...

Juan Carlos Munoz / Jersey tiger moth (Euplagia quadripunctaria) large numbers flying to and resting on Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011

MYN / Marko Masterl / Viper's bugloss (Echium vulgare) in flower with solitary bee (Panurgus sp), Slovenia, Europe, June. meetyourneighbours.net project

Martin Gabriel / Four Griffon Vultures (Gyps fulvus) in flight, silhouetted overhead. Monfrague National Park, Extremadura, Spain, March.

John Abbott / Ichneumon wasp (Compsocryptus sp.?) in flight, Williamson county, Texas, USA, November

John Abbott / Composite image of flying insects visiting blacklight, Texas, USA, September