Juergen Freund / Fan coral (Gorgonacea) in the shallows, surrounded by mangrove roots and tiny fish. North Raja Ampat, West Papua, Indonesia, February.
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Andy Rouse / Polar Bear (Ursus maritimus) portrait in sea-ice landscape. Wide angle / fish-eye shot. Svalbard, Norway, September.
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Doug Perrine / Bioluminescence from glowing plankton in sea tide line on beach, with stars above and ship lights on horizon, Vaadhoo Island, Raa Atoll, Maldives, Indian Ocean, October 2010
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Sergio Hanquet / Green sea turtle (Chelonia mydas), two viewed from above. Tenerife, Canary Islands.
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Danny Green / Polar bear (Ursus maritimus) resting on sea ice, view from above. Svalbard, Norway, June 2018.
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Guy Edwardes / Milky Way above Durdle Door rock arch. Jurasic Coast, Dorset, England, UK. August 2019. Composite image.
Alex Mustard / School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean
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Alex Mustard / Veined octopus (Amphioctopus marginatus) shelters inside an old clam shell, with silver-sides in the water above. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Espen Bergersen / Aerial shot of Killer whale (Orcinus orca) hunting Atlantic herring (Clupea harengus) using carousel feeding, with herring jumping out of the water and gulls (Larus) circling above, Tromvik, Troms, No...
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Marie Read / Horned Puffin (Fratercula corniculata) flying over a pattern of kelp and sea foam, photographed from above, St. Paul, Pribilof Islands, Alaska, USA. July.
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George Sanker / North American river otter (Lontra canadensis) swimming with head above water. Acadia National Park, Maine, USA. November.
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Sven Zacek / Wild boar (Sus scrofa) swimming with head above water in Suur Emajogi River. Alam-Pedja Nature Reserve, Tartumaa, Southern Estonia. November.
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Nick Hawkins / Galapagos green turtle (Chelonia mydas agassizi) swimming above sea floor with shoal of fish in background. Off San Cristobal Island, Galapagos.
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Jack Dykinga / Aurora Borealis display above conifers and Polar Lake. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
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David Fleetham / Whale shark (Rhincodon typus), a commercial encounter for tourists with feeder above on canoe. Oslob, Cebu, Philippines. April 2017.
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Alex Mustard / Hawksbill turtle (Eretmochelys imbricata) pair circling each other above coral reef. Seven Mile Beach, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
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Theo Bosboom / Edible frog (Pelophylax k. esculentus) catching a common blue damselfly (Enallagma cyathigerum) with its tongue with pair mating on stem above, Hondenven, Tubbergen, The Netherlands, July.
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SCOTLAND: The Big Picture / Pod of Bottlenose dolphins (Tursiops truncatus) surfacing just offshore, Black Isle, Scotland, UK, July.
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Ingo Arndt / European edible frog eyes showing above water {Rana esculenta} Germany
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Christophe Courteau / African elephants (Loxodonta africana) swimming across Chobe River, trunks raised above surface for breathing, Chobe National Park, border between Botswana and Namibia.
Alex Mustard / Grey seal (Halichoerus grypus) playful young seal above a kelp forest. Lundy Island, Devon, UK Bristol Channel, August
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Claudio Contreras / American crocodile (Crocodylus acutus) resting just above seagrass underwater, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
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Alex Mustard / Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
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Tony Wu / Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean.
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Floris van Breugel / Temple Crag above Second lake, one the of the Big Pine lakes below the Palisade basin, John Muir Wilderness, Sierra Nevada, California, USA, May
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Bence Mate / Great egrets (Ardea alba) in winter, and Grey heron (Ardea cinerea) with flock of other birds flying above, Lake Csaj, Kiskunsagi National Park, Hungary. January.
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Espen Bergersen / Gulls (Laridae) seaching for food in water containing blood coming from a large fishing boat, Tromvik, Kvaloya, Troms, Norway, November 2014.
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Wild Wonders of Europe / Widstrand / Humpback whale (Megaptera novaeangliae) tail fluke above water before diving, Senja, Troms County, Norway, Scandinavia, January. Cetaceans are attracted to this area to feed on the large numbers of sp...
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Wild Wonders of Europe / Widstrand / Humpback whale (Megaptera novaeangliae) tail fluke above water before diving, Senja, Troms County, Norway, Scandinavia, January. Cetaceans are attracted to this area to feed on the large numbers of sp...
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Pedro Narra / Hippopotamus (Hippopotamus amphibius) with head raised above water surface, Bijagos Archipelago Biosphere Reserve, Guinea Bissau,
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Alex Mustard / Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
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Alex Mustard / Hawksbill turtle (Eretmochelys imbricata) male swimming in open water above a coral reef. Tank Rock, Fiabacet, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea, Tropical West Pacific Ocean.
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Mark MacEwen / Japanese firefly (Luciola cruciata) light trails at night above stream in rural Japan. June.
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Olga Kamenskaya / Clouds over Lake Baikal, Siberia, Russia, October 2010.
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George Sanker / North American river otter (Lontra canadensis) two in water with autumnal trees reflected in the water, Acadia National Park, Maine, USA, October.
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David Tipling / Common frog (Rana temporaria) and spawn in pond. West Runton, North Norfolk, England, UK, March. Highly commended in the Habitat category of the BWPA Awards 2015.
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Chris Mattison / Budgett's Frog (Lepidobatrachus laevis) mostly submerged in muddy water, with eyes visible above water, captive, occurs in South America.
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Ole Jorgen Liodden / Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.
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Steven Kazlowski / Photographer in front of Northern lights / Aurora borealis glowing brightly over the frozen eastern Beaufort Sea, Arctic National Wildlife Refuge, Alaska Model released.
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Espen Bergersen / Humpback whale (Megaptera novaeangliae) with fin above water, Andfjorden close to Andoya, Nordland, Northern Norway. January.
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Kirkendall-Spring / Mount Rainier reflected in tarn above Mystic Lake in Mount Rainier National Park, Washington, USA, August 2014.
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Pascal Kobeh / Female humpback whale (Megaptera novaeangliae) with its very young calf at the surface, Rurutu island, Australs Archipelago, French Polynesia. Pacific Ocean.
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Hougaard Malan / Tugela falls surrounded by clouds on a misty morning. Royal Natal National Park, Drakensberg, South Africa. November 2010.
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Alex Mustard / World War II British motorbikes (Norton 16H) stacked up on the back of truck in the hold of wreck of HMS Thistlegorm, with soldierfish (Myripristis murdjan) swimming above. Sha'ab Ali, Sinai, Egypt. R...
Andy Trowbridge / Grey Seal (Halichoerus grypus) at rest on submerged rock, head and one flipper above water. Shetland Islands, Scotland, UK. July.
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Andy Trowbridge / Common / Harbour seal (Phoca vitulina) looking out of water with bee hovering above its nose. Shetland Islands, Scotland, UK, July.
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Jussi Murtosaari / Daubenton's bat (Myotis daubentonii) hunting a mosquito. central Finland, June.
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Tim Laman / Aerial view of morning mist above the Tiputini River in Yasuni National Park, Francisco de Orellana Province, Ecuador, July.
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Chris & Monique Fallows / Split level shot of Great white shark (Carcharodon carcharias) fin above the water and leisure boat in the background, with Seal Island, False Bay, South Africa.
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Alex Mustard,Alex Mustard / Green surf anemones (Anthopleura xanthogrammica) grow in shallow water below the temperate rain forest. Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada. North East Pacific Ocean.
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Laurent Geslin / Female Emperor dragonfly (Anax imperator) close-up, on twig above water with clouds reflected, Europe
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Theo Bosboom,Theo Bosboom / Gulls, predominantly Iceland gulls (Larus glaucoides) flying above stormy ocean near Vik, Iceland, March 2012.
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Theo Bosboom / Fulmar (Fulmarus glacialis) above snowy lava beach near Djupivogur in the East of Iceland, March
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Felicity Lanchester,Felicity Lanchester / Cloud iridescence forming above a cumulonimbus cloud, caused by light refraction through individual water droplets and ice crystals at high altitude, Sierra Leone.
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Erlend Haarberg,Erlend Haarberg / Aerial view down river to the east of the Hofsjakull glacier and west of the Sprengisandur upland desert. Innumerable rivulets flow from beneath the glacier in this area of tundra which abounds in pon...
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Franco Banfi / Great Blue shark (Prionace glauca) profile portrait viewed from slightly above, Pico Island, Azores, Portugal, Atlantic Ocean
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Andy Trowbridge / White-tailed sea eagle (Haliaeetus albicilla) in flight. Flatanger, Norway, May.
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Michel Roggo / Grayling (Thymallus thymallus) on spawning ground. Lac de Fenetre, alpine lake, 2456 m above sea level, Canton of Valais, Switzerland, June.
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Sandra Bartocha / Dark clouds above the North Sea, Norderney, Germany, September
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Brent Stephenson / Flying fish (probably Cypselurus lineatus) in flight above the water. Off Whitianga, Coromandel Peninsula, New Zealand.
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Paul Williams / Tourists watching two wild Jaguar cubs fight on riverbank whilst third looks on from above (Panthera onca) from boat on river, Pantanal, Pocone, Brazil. No release available.
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Roy Mangersnes / Polar bear (Ursus maritimus) holding piece of ice after licking off freshwater, Spitsbergen, Svalbard, Norway, August
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Nick Turner / Crater lake with reflections of clouds in the water, Queen Elizabeth National Park, Uganda, January 2009.
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Alex Mustard / Spotted porcupinefish (Diodon hystrix) swimming in open water above a coral reef, West Bay, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.
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Graham Eaton / View towards Bardsey Island / Ynys Enlli, Aberdaron, Gwynedd, Wales, UK, January
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Inaki Relanzon / Whitsunday Islands, aerial view, Great Barrier Coral Reef, Queensland, Australia, October 2011.
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Inaki Relanzon / Whitsunday Islands, aerial view with hydroplane landing, Great Barrier Coral Reef, Queensland, Australia, October 2011.
LYNN M. STONE / Bullfrog (Rana catesbeiana) with head sticking out above water in pond, Deep River, Connecticut, USA, July
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Ben Osborne / Aerial view of African elephants (Loxodonta africana)moving through wetland in the Okavango delta, Northern Botswana, taken on location for BBC Planet Earth series, October 2005
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Oriol Alamany / People and an iceberg in Lake Grey with Valle del Frances in the background, Torres del Paine National Park, Patagonia, Chile
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Sven Zacek / River meeting the sea on the Baltic coast of Estonia. Europe, May 2011.
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Sven Zacek / Lowland marshes seen from the air. Estonia, Europe, October 2010.
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Sven Zacek / Lowland bog landscape seen from the air. South Estonia, Europe, October 2010.
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Andy Rouse / Polar ice cap at sunset. Austfonna Polar Ice Cap, Svalbard, Norway, August 2011.
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Andy Rouse / 2020VISION / European river otter (Lutra lutra) swimming with head above water, river, Dorset, UK, November
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Martin Dohrn / Starlight camera image in moonlight of Eurasian wild boar (Sus scrofa) feeding on carcass of Spotted deer stag (Axis axis) watched by Asian water buffalo (Bubalas bubalas) while the leopard (Panthera...
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Robert Thompson / Lighthouse on Fanad Head at sunset, County Donegal, Republic of Ireland, UK, August 2011
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Ole Jorgen Liodden / Polar Bear (Ursus maritimus) sitting on floe in vast thawing sea-ice landscape. Spitsbergen, Svalbard, August 2011. Second place, Mammals Category, GDT 2012 competion. Winner in ANIMALS IN THEIR ENVIR...
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Ole Jorgen Liodden / Polar Bear (Ursus maritimus) looking across ice floe seascape. Nordaustlandet, Svalbard, August.
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Ole Jorgen Liodden / Blue Whale (Balaenoptera musculus) tail fluke above water surface. Spitsbergen, Svalbard, August.
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Kerstin Hinze / Aerial view of Westerhever Sand lighthouse and saltmarshes, Wadden Sea, Eiderstedt, Schleswig Holstein, Germany, June 2010
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Jerry Monkman / Lake Gloriette and the Balsams Grand Resort as seen from the cliffs above NH 26 in Dixville Notch, New Hampshire, USA, September 2009
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Joe Cornish / 2020VISION / Pool and Stac Pollaidh, Coigach / Assynt SWT, Sutherland, Highlands, Scotland, UK, June 2011. Did you know? The peak of Stac Polliadh is very severely weathered and it is thought that during the last...
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Orsolya Haarberg / Red Knots (Calidris canutus) during migration rest on coastal rocks as waves wash across the coastline. The Snæfellsnes peninsula, Iceland, May. Highly commended in the Birds category of the GDT 2011...
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Bryan and Cherry Alexander / Meltwater pouring off iceberg above hole. Scoresbysund Fiord, East Greenland, September 2005.
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Mark Hamblin / 2020VISION / Rowan tree silhouetted above Loch Lurgainn with Cul Mor (left) and Ben More Coigach beyond, Coigach, Highland, Scotland, UK, November. Photographer quote: "Let's face it walking uphill is a drag – I g...
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Hermann Brehm / Young Tigress (Panthera tigris), 22 month, being frightened by a dead leaf. Banghavgarh National Park, India, April.
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Eric Baccega / Narwhal (Monodon monoceros) showing tusk above water surface. Baffin Island, Nunavut, Canada, April.
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Eric Baccega / Narwhal (Monodon monoceros) showing tusk above water surface. Baffin Island, Nunavut, Canada, June.
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Eric Baccega / Narwhal (Monodon monoceros) showing tusks above water surface. Baffin Island, Nunavut, Canada, June.
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Eric Baccega / Narwhal (Monodon monoceros) crossing tusks above water surface. Baffin Island, Nunavut, Canada, April.
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Inaki Relanzon / Isolated rain storm above the sea. Bahamas Sea, Bahamas, North America, August 2006.
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Inaki Relanzon / Aerial view of the mangroves around Betsiboka River. Majunga, north-west Madagascar, Africa, October 2008.