Rick Tomlinson / Farr 47 "Ausmaid" battles upwind during the Sydney-Hobart Race, 1997. ^^^Finishing 3rd overall.

Benjamin Barthelemy,Benjamin Barthelemy / Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June

Nature Production / Bombardier Beetle (Pheropsophus jessoensis) protecting itself by ejecting a boiling, noxious chemical spray, Nagasaki, Japan.

Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition

Morley Read / Slow motion clip of a Linnaeus' sipo snake (Chironius exoletus) with open mouth, part of threat display, Amazon rainforest, Orellana Province, Ecuador.

Morley Read / Parrot snake (Leptophis ahaetulla) threat display, Amazon rainforest, Orellana Province, Ecuador.

Bence Mate / European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition.

Diane McAllister / Polar bears (Ursus maritimus) fighting in water. Beaufort Sea , Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Nature's Best Photography Competition 2019.

David Tipling / Common European adder (Vipera berus) two males fighting over a female. Holt, North Norfolk, England, UK. April

David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April

David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.

Alex Mustard / Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...

Tony Wu / View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.

Tony Wu / Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tony Wu / Zoarchias major eelpouts. (Zoarchias major) The orange individual is female, the one in the hole is male. The female has approached the male, curled around him, attempting to dislodge the male from th...

Tony Wu / Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.

Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.

Nick Hawkins / European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July.

John Chan / Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...

Marie Read / Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October.

Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA

Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.

David Pattyn / Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May

Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.

Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.

George Sanker / Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.

Eric Baccega / Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia

Staffan Widstrand / Wild Wonders of China / Pere David's deer / Milu (Elaphurus davidianus), two stags testing their strength in Hubei Tian'ezhou Milu National Nature Reserve, Shishou, Hubei, China

Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults brooding chicks. One chick exhibiting aggression towards other. Atka Bay, Antarctica. September.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.

Guy Edwardes / Mozambique spitting cobra (Naja mossambica) spitting venom. Kruger, South Africa. Controlled conditions.

Paul Hobson / Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica.

Paul Hobson / Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.

Pal Hermansen / Brown bear (Ursus arctos), two showing aggression at night. Finland. August.

David Perpinan / Group of Griffon vultures (Gyps fulvus) fighting to feed on a dead lamb. Cuenca, Spain, August.

Andy Rouse / White-necked Jacobin hummingbirds (Florisuga mellivora) fighting, Tobago Small repro only

Kerstin Hinze / Two Black-bellied hamsters (Cricetus cricetus) fighting, Vienna, Austria, October.

Franco Banfi / Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica.

Tui De Roy / San Cristobal giant tortoises (Chelonoidis chatamensis), fighting, Galapaguera, San Cristobal Island, Galapagos

Philippe Clement / Two Lynx (Lynx lynx) kittens play fighting in front of den entrance, Bavarian Forest National Park, Germany, October. Captive.

David Fleetham / Great white shark (Carcharodon carcharias) breaking surface with mouth open. Guadalupe Island, Mexico.

Remi Masson / Pair of Pumpkinseeds (Lepomis gibbosus) mating, male chasing other individuals away, River Rhone, Rhone-Alpes, France, June.

Remi Masson / Male Common toads (Bufo bufo) fighting underwater during mating season, with a pair mating Ain, Rhone-Alps, France, March.

Dave Bevan / Slow motion clip of two pairs of Jackdaws (Corvus monedula) squabbling over nest site, Worcestershire, England, UK, May.

David Fleetham / Jaws and teeth of Stout Moray (Gymnothorax eurostus). Hawaii.

Sergey Gorshkov / Northern goshawk (Accipiter gentilis) flying with squirrel prey, Finland . March

Sergey Gorshkov / Northern goshawks (Accipiter gentilis) fighting over squirrel carcass, Finland, March.

Sergey Gorshkov / Black Grouse (Tetrao tetrix) males fighting at lek in the snow, Tver, Russia. April

Five Films / Female Grey seal (Halichoerus grypus) defending pup on beach, chasing male away, Horsey, Norfolk, November.

Dave Bevan / Juvenile male Wild boar (Sus scrofa) fighting to establish seniority, Gloucestershire, England, UK, December.

Philippe Clement / Red deer (Cervus elaphus) stags fighting, Scotland, UK, May.

Erlend Haarberg / Mountain hares (Lepus timidus) fighting in snow at night, Vauldalen, Norway. Joint overall winner of the GDT European Wildlife Photographer of the Year 2017. Highly commended in the Wildlife Photo...

Frederique Olivier / Group of Adelie penguins (Pygoscelis adeliae) destroying an egg with a chick inside, Adelie Land, Antarctica, January.

Brian Bevan / Brown hare (Lepus europaeus) boxing, Bedfordshire, England, UK, March.

Dave Bevan / Two male Grey seals (Halichoerus grypus) fighting in surf, with a female and juvenile nearby, Norfolk, England, UK. December.

Francisco Marquez / Two male Iberian wild goats (Capra pyrenaica) fighting, Sierra de Gredos, Spain, November.

Sandesh Kadur / Two Bengal monitors (Varanus bengalensis) fighting, Ranthambore National Park, Rajasthan, India. 2016.

Philippe Clement / Two male Alpine ibex (Capra ibex) fighting on a steep mountain rock face, Gran Paradiso National Park, Graian Alps, Italy, June.

Frederique Olivier / Chinstrap penguin (Pygoscelis antarcticus) adding a stick to its nest and fighting with a neighbouring pair, Aitcho Island, South Shetland Islands.

Nick Upton / Rock goby (Gobius paganellus) stealing food from a scavenging Common prawn (Palaemon serratus) in a rockpool, Cornwall, England, UK, September.

Sandesh Kadur / Two male King cobras (Ophiophagus hannah) fighting, Agumbe, Karnartaka, India.

David Pattyn / Black necked grebe (Podiceps nigricollis) pair chasing away another grebe during the courtship season. The Netherlands.April 2014

Ammonite / Male Yellow dung flies (Scathophaga stercoraria) fighting over a female on a cowpat, UK.

Ammonite / Male Yellow dung flies (Scathophaga stercoraria) fighting over a female on a cowpat, UK.

Dietmar Nill / Two European hares (Lepus europaeus) boxing in a snow covered field, Germany, March.

Konrad Wothe / Showreel of Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.

Ingo Arndt / Blue Crab (Discoplax hirtipes) with claws raised in defensive / aggressive posture, endemic to Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Blue Crab (Discoplax hirtipes) with claws raised in defensive / aggressive posture, endemic to Christmas Island, Indian Ocean, Australian Territory

Ingo Arndt / Eastern grey kangaroos sparring {Macropus giganteus} Australia

Ingo Arndt / Two Northern elephant seal (Mirounga angustirostris) males fighting for dominance, California, USA

Ingo Arndt / Hamadryas baboon male threat display {Papio hamadryas} Africa

Ingo Arndt / Jackson's chameleons fighting {Chamaeleo jacksonii} Kenya

Ingo Arndt / Gelada baboon {Theropithecus gelada} juveniles playing on rocks, Simien Mountains NP, Ethiopia

Ingo Arndt / Gelada baboon males threat display {Theropithecus gelada} Simien Mt NP, Ethiopia

Ingo Arndt / Gelada baboon male threat display {Theropithecus gelada} Simien Mt NP, Ethiopia

Konrad Wothe / Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.

SCOTLAND: The Big Picture / Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April.

SCOTLAND: The Big Picture / Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April.

Ingo Arndt / Masai giraffe (Giraffa camelopardalis tippelskirchi) males fighting, Masai Mara National Reserve, Kenya. Sequence 3 of 3

Carol Walker / Two wild Mustang stallions fighting in the White Mountain Herd Area, Wyoming, USA. August.

Sergey Gorshkov / Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa.

Jordi Chias / Two male Dusky groupers (Epinephelus marginatus) fighting for territory during mating season at 'baixa do meros', a small pinnacle at Formigas islets Protected area, near Santa Maria Island, Azores. F...

Neil Aldridge / Plain's zebra (Equus quagga) biting the neck of a rival during a fight Hluhluwe imfolozi Park. South Africa.

Neil Aldridge / Hippopotamus (Hippopotamus amphibius) charging through the shallows, Chobe River, Chobe National Park, Botswana. Vulnerable species.

Danny Green / Coastal brown bears (Ursus arctos) fighting, Lake Clarke National Park, Alaska, September

Will Burrard-Lucas / Aardvark (Orycteropus afer) fighting in the dark. Makgadikgadi Pans, Botswana. January

Yashpal Rathore / Asiatic elephant (Elephas maximus), female charging, Jim Corbett National Park, India. 2014

Bence Mate / Great egrets (Ardea alba) fighting with necks intertwined, Lake Csaj, Kiskunsagi National Park, Hungary. January.

Bence Mate / Juvenile White tailed sea eagles (Haliaeetus albicilla) fighting for fish in mid air, Lake Csaj, Kiskunsagi National Park, Hungary. February.

Bence Mate / White tailed sea eagle (Haliaeetus albicilla) interacting with Great egret (Ardea alba) Lake Csaj, Kiskunsagi National Park, Hungary. January.

Bence Mate / Two Buzzards (Buteo buteo) fighting, Lake Csaj, Kiskunsagi National Park, Hungary. February.

Tony Heald / African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana

Bence Mate / Great white egret (Ardea alba) fighting with Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Hungary, January.

Bence Mate / Caspian gulls (Larus cachinnans) juveniles fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. March.