Ann & Steve Toon / Impala (Aepyceros melampus) allo-grooming, iMfolozi game reserve, KwaZulu-Natal, South Africa, August
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Anup Shah / Western chimpanzee (Pan troglodytes verus) young male 'Jeje' aged 13 years being groomed by young male 'Peley' aged 12 years, Bossou Forest, Mont Nimba, Guinea. January 2011.
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Anup Shah / Western chimpanzee (Pan troglodytes verus) males 'Jeje' aged 13 years, 'Tua' aged 53 years and 'Peley' aged 12 years grooming, Bossou Forest, Mont Nimba, Guinea. January 2011.
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Chris Gomersall / King penguins (Aptenodytes patagonicus) allo-preening. Fortuna Bay, South Georgia, Antarctica.
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Mark Carwardine / Yellow-eyed penguins / Hoiho (Megadyptes antipodes) allo preening one another, Otago Peninsula, South Island, New Zealand, June, endangered species
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David Tipling / Wandering albatross (Diomedea exulans) pair allo grooming, part of ritual bonding behaviour, Albatross Island, South Georgia, January
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David Tipling / Wandering albatross (Diomedea exulans) pair allo preening on Albatross Island, South Georgia, Januar
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David Tipling / Wandering albatross (Diomedea exulans) pair allo preening on Albatross Island, South Georgia, January
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Richard Du Toit / Burchell's zebra (Equus quagga burchellii) allo grooming, Gauteng Province, Rietvlei Nature Reserve, South Africa.
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David Tipling / Eurasian Coots (Fulica atra allo) preening on Lake Geneva, Switzerland, March.
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Anup Shah / Western chimpanzee (Pan troglodytes verus) young male 'Jeje' aged 13 years grooming young male 'Peley' aged 12 years, Bossou Forest, Mont Nimba, Guinea. January 2011.
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Anup Shah / Western chimpanzee (Pan troglodytes verus) males 'Jeje' aged 13 years, 'Tua' aged 53 years and 'Peley' aged 12 years grooming, Bossou Forest, Mont Nimba, Guinea. January 2011.