Jussi Murtosaari / Arran brown butterfly (Erebia ligea) feeding on Wild angelica (Angelica sylvestris), Jyvaskyla, Finland, July

Nick Upton / Club-horned sawfly (Abia sericea) feeding on Wild angelica (Angelica sylvestris) flowers, chalk grassland meadow, Wiltshire, UK, July.

Nick Upton / Buff-tailed Bumblebee (Bombus lucorum) foraging on Wild Angelica (Angelica sylvestris) flowering on river bank. Wiltshire, UK, June.

Nick Upton / Green sawfly (Rhogogaster viridis) feeding on Wild Angelica (Angelica sylvestris) flowers on river bank, Wiltshire, UK, June.

Nick Upton / Male hoverfly (Syritta pipiens) feeding on Wild Angelica (Angelica sylvestris) flowerhead on river bank. Wiltshire, UK, June.

Nick Upton / A large black and yellow Ichneumon Wasp (Amblyteles armatorius), a parasitoid of noctuid moth caterpillars, feeding on Wild Angelica (Angelica sylvestris) flowers. Wiltshire river bank, UK, June.

Nick Upton / Moth caterpillar (Depressaria daucella) a specialist of riverine habitats, stretching up from one cluster of Angelica (Angelica sylvestris) flowers to another, Wiltshire riverbank, UK, June.

Nick Upton / Moth caterpillar (Depressaria daucella) a specialist of riverine habitats, feeding on Angelica (Angelica sylvestris) flowers, Wiltshire riverbank, UK, June.

Nick Upton / Moth caterpillar (Depressaria daucella) a specialist of riverine habitats, feeding on Angelica (Angelica sylvestris) flowers, Wiltshire riverbank, UK, June.

Nick Upton / White form of Goldenrod crab spider (Misumenia vatia) camouflaged on Wild Angelica (Angelica sylvestris) flowers sinking fangs into the head of a large Horse fly (Dasyrhamphis anthracinus). Corsica, F...

Nick Upton / White form of Goldenrod crab spider (Misumenia vatia) camouflaged on Wild Angelica (Angelica sylvestris) flowers sinking fangs into the head of a large Horse fly (Dasyrhamphis anthracinus). Corsica, F...

Nick Upton / Male Hoverfly (Myathropa florea) feeding on Wild angelica (Angelica sylvestris) umbel flowers. Corsica, France, June.