Martin Dohrn / Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using 'Starlight Camera' technology without artificial lighting.

Alex Mustard / Synchronous spawning of a mountainous star coral (Montastraea faveolata) at night. East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.

Richard Du Toit / Burchell's Zebra (Equus burchellii) pair nuzzling, Rietvlei Nature Reserve, Gauteng, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) yawning, Rietvlei Nature Reserve, Gauteng, South Africa

Richard Du Toit / Cattle Egret (Bubulcus ibis) on Burchell's Zebra (Equus burchellii), Rietvlei Nature Reserve, Gauteng, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) pair, Rietvlei Nature Reserve, Gauteng, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa

XI ZHINONG / Grey wolf (Canis lupus) chasing a wounded tibetan wild ass (Equus kiang) Kekexili, Qinghai, China, December.

XI ZHINONG / Grey Wolves (Canis lupus) hunting a wounded Tibetan wild ass (Equus kiang) Kekexili, Qinghai, China, December.

Alex Mustard / A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea.

Vladimir Medvedev / Camera trap image of wild Siberian tiger (Panthera tigris altaica) walking through woodland, Lazovsky Zapovednik Nature Reserve, Primorsky Krai, Far East Russia, November 1994

Konrad Wothe / Prehistoric Rock Art, Awis Valley, Akakus mountains, Libya, Sahara, North Africa, November 2007

Konrad Wothe / Stone engravings of Giraffes in Wadi Mathendous, Wadi Barjuj, Stony Desert, Libya, Sahara, North Africa, November 2007

Erlend Haarberg / Rock ptarmigan (Lagopus muta) in snow near glacier, Iceland, August 2010

Ben Hall / European rabbits (Oryctolagus cuniculus) juveniles emerging from burrow, Cheshire, UK May

Vladimir Medvedev / Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species. Commended in Wildlife Photograp...

Florian Möllers / Quinling Golden snub nosed monkey (Rhinopitecus roxellana qinlingensis), family group resting and grooming in a stand of trees. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China.

Andy Sands / Digger Wasp (Cerceris arenaria) carrying large weevil back to burrow, paralysed as food for larvae. London, England, UK, July.

Marie Read / Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February

Roy Mangersnes / Black-legged kittiwakes (Rissa tridactyla) in flight during evening snowfall, Northern Nordaustlandet, Svalbard, Norway, August

Roy Mangersnes / Wild Bolivian / Peruvian Squirrel Monkey (Saimiri boliviensis) in tree. Madidi National Park, Bolivia.

Roy Mangersnes / Arctic fox (Alopex / Vulpes lagopus) in tundra landscape, near Longyearbyen, Spitsbergen, Svalbard, Norway, August

Roy Mangersnes / Arctic fox (Alopex / Vulpes lagopus) standing, near Longyearbyen, Spitsbergen, Svalbard, Norway, August

Roy Mangersnes / Two Barnacle geese (Branta leucopsis) in flight, Spitsebergen, Svalbard, Norway, August

Gerrit Vyn / Spotted owl (Strix nebulosa) Gifford-Pinchot National Forest, Washington USA

Paul Hobson / Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK.

Markus Varesvuo / Barn swallow feeding chick on the wing (Hirundo rustica) Sipoo Finland July

Mary McDonald / Hanuman Langur / Northern Plains Grey Langur (Semnopithecus entellus) in leafless tree top. Kanha National Park, Madhya Pradesh, India.

John Abbott / Tourists watching large colony of Marine Iguanas (Amblyrhynchus cristatus) Punta Espinosa on Fernandina Island, Galapagos, August 2010

Juan Carlos Munoz / Jersey tiger moth (Euplagia quadripunctaria) large numbers resting on leaves of Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011

Juan Carlos Munoz / Jersey tiger moth (Euplagia quadripunctaria) large numbers resting on trunk of Sweetgum tree (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011

Juan Carlos Munoz / Jersey tiger moth (Euplagia quadripunctaria) large numbers resting on leaves of Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011

Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on. Scandinavia, April. 5th, 2010 Euronatur Wildlife photographer competition.

Juan Carlos Munoz / White Tailed Sea Eagles (Aliaeetus albicilla) flying above snow with a Raven (Corvus corax) landed. Scandinavia, Europe, April.

Sven Zacek / Geese on their spring migration above dawn lit clouds. Estonia, Europe, May.

Sven Zacek / Roe Deer (Capreolus capreolus) males with developing antlers in velvet, one in focus, one not. Virumaa, Estonia, Europe, March. Winner of the Animal stories portfolio in the Melvita Nature Images Awar...

Sven Zacek / Roe Deer (Capreolus capreolus) from different angles, only one in focus. Virumaa, Estonia, Europe, March. Winner of the Mammals category in GDT competition 2013. 2nd Prize in the Mammals Category of M...

Sven Zacek / Fish seen through water and the reflections of trees. Jogeva maakond, Estonia, Europe, February.

Sven Zacek / Large herd of Roe Deer (Caperolus capreolus) running across snow. Virumaa, Estonia, Europe, March.

Ole Jorgen Liodden / Blue Whale (Balaenoptera musculus) blowing, with Black-legged Kittiwake (Rissa tridactyla) in the water spray. Spitsbergen, Svalbard, August. Runner up in the Birds category of GDT competition 2014.

Sandesh Kadur / Termites (Isoptera) flying upwards towards a street lamp, India

B&S Draker / Pipevine Swallowtail Butterfly (Battus philenor) pair mating. Sinton, Corpus Christi, Coastal Bend, Texas, USA, June.

B&S Draker / Cicada (Tibicen resh) adult emerging from nymph carapace. Sinton, Corpus Christi, Coastal Bend, Texas, USA, June. Sequence 10 of 10.

Bence Mate / Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competit...

Sven Zacek / Roe deer (Capreolus capreolus) in foreground with second deer on horizon in snow, Estonia, December. Highly commened in CREATIVE VISIONS OF NATURE category, 2011 WILDLIFE PHOTOGRAPHER OF THE YEAR COM...

Terry Whittaker / 2020VISION / Grey Squirrel (Sciurus carolinensis) climbing over fence in parkland, Regent's Park, London, UK, April 2011. Photographer quote: 'Grey squirrels may be overfed, oversexed and over here, but they are s...

Georgette Douwma / Barrel Sponge (Xestospongia testudinaria) on coral reef with soft corals (Scleronephthya sp), gorgonian and Lyretail anthias (Pseudanthias squamipinnis). Komodo National Park, Indonesia.

Pete Oxford / Red Mangrove (Rhizophora mangle) and fish. Sian Ka'an Biosphere Reserve, Quintana Roo, Yucatan Peninsula, Mexico.

Juergen Freund / Green turtles (Chelonia mydas) mating over coral reef, Sipadan Island, Sabah, Malaysia, June, Endangered species

Juergen Freund / Schooling Silverside fish (Antherinidae) split level with karst limestone walls, fan coral and featherstar, North Raja Ampat, West Papua, Indonesia, February 2002.

Steve Nicholls / Huge flock of Common starlings (Sturnus vulgaris) flying in to roost over the reed beds at dusk, Ham Wall nature reserve, Somerset, UK, January

Juergen Freund / Crinoids or featherstars covering a coral bommie with dive boat looming overhead, West New Britain, Papua New Guinea

Andy Rouse / Portrait of a Mountain Gorilla (Gorilla beringei) with a view over forest canopy. Rwanda, Africa, March.

Juergen Freund / Schooling Yellowtail fusilier (Caesio cuning) Misool, Raja Ampat, West Papua, Indonesia

Juergen Freund / Schooling Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia

Juergen Freund / Schooling Banggai cardinalfish in anemone tentacles. (Pterapogon kaudemi). Lembeh Strait, North Sulawesi, Indonesia

Juergen Freund / Snorkeler swimming through a school of Oxeye scads (Selar boops) in the house reef of Miniloc Island Resort, El Nido island, Palawan, Philippines, May 2009. Model released.

Juergen Freund / Giant trevally (Caranx ignobilis) chasing shoal of Oxeye Scads (Selar boops) in the house reef of Miniloc Island Resort, El Nido, Palawan, Philippines

Andy Rouse / Solitary Grizzly Bear (Ursus arctos horribilis) in expansive mountain landscape. Katmai, Alaska, USA, August.

Andy Rouse / Grizzly Bear (Ursus arctos horribilis) walking through a river with caught salmon against a mountainous landscape as two gulls fly past. Katmai, Alaska, USA, August.

Andy Rouse / Grey Wolf (Canis lupus) with clouds and a glacier in the distance. Katmai, Alaska, USA, August.

Juergen Freund / Newly hatched Leatherback turtle (Dermochelys coriacea) babies moving from the nest towards the sea. Warmamedi beach, Bird's Head Peninsula, West Papua, Indonesia, July 2009

Juergen Freund / Yellow-ribbon sweetlips (Plectorhinchus polytaenia) surrounded by dense shoal of glassy sweepers (Pempheris schomburgki). North Raja Ampat, West Papua, Indonesia

Tony Phelps / Southern Dwarf Adder (Bitis armata) climbing in dune thicket. De Hoop Nature Reserve, Western Cape, South Africa, January.

Tony Phelps / Scorpion (Uroplectes lineatus / carinatus) female carrying hatchlings on its back; a rare example of parental care in invertebrates. De Hoop Nature Reserve, Western Cape, South Africa, January.

Edwin Giesbers / Konik Wild Horse (Equus ferus caballus) in grasslands with Purple loosestrife flowers (Lythrum salicaria) in foreground. The Netherlands, July.

Chien Lee / Spectral Tarsier (Tarsius tarsier), Tangkoko Nature Reserve, Sulawesi, Indonesia

Orsolya Haarberg / Spotted redshank (Tringa erythropus) at shallow water in snow, Ovre-Pasvik National Park, Norway

David Noton / James' flamingos (Phoenicoparrus jamesi) on Lake Colorada in the remote region of high desert, altiplano and volcanoes near Tapaquilcha, Bolivia, December 2009

Chien Lee / Lilac-cheeked Kingfisher (Cittura cyanotis), Tangkoko Nature Reserve, Sulawesi, Indonesia

Konstantin Mikhailov / West Caucasian tur (Capra caucasica) Female on cliff face, Highlands of West Caucasus Mountains in the Bezengi area, Kabardino-Balkaria Republic, Kabardino-Balkarsky State Nature Reserve, Russia.

Markus Varesvuo / Bearded vulture (Gypaetus barbatus) flying, Spain, November

Markus Varesvuo / Bearded vulture (Gypaetus barbatus) flying, Spain, November

Kristel Richard / Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa, O...

Barrie Britton / Lesser flamingos (Phoenicopterus minor). Lake Bogoria, Kenya. Picture taken during filming for BBC "Life" TV Series, May 2008

Bernard Castelein / Cinereous / European Black Vulture (Aegypius monachus) in dispute with an Eurasian Griffon Vulture (Gyps fulvus), alongside a Himalayan Griffon (Gyps himalayensis) and Long-billed Vulture (Gyps indicu...

Jenny E. Ross / Snow Geese (Chen caerulescens) and Ross's Geese (Chen rossii) taking off from field where they have been feeding, Imperial Valley, California, USA.

Eric Baccega / Moose (Alces alces) bull walking in forest clearing, Cap Breton Highlands National Park, Nova Scotia, Canada, September

Charlie Summers / King Penguin (Aptenodytes patagonicus) adult surrounded by huddled chicks, riding out a snowstorm at Gold Harbor, South Georgia.

Charlie Summers / King Penguin (Aptenodytes patagonicus) adult standing amongst colony of chicks, Gold Harbor, South Georgia, South Atlantic Islands

Charlie Summers / Two Mustang (Equus caballus) stallions, a strawberry roan on the left and a black on the right, rearing and fighting for dominance, Pryor Mountain Range of Montana, USA

Kevin Schafer / Peruvian pelicans (Pelecanus thagus) on coastal rocks, Pucusana, Lima Province, Peru

Kevin Schafer / Kittiwake (Rissa tridactyla) flock feeding in from of the Monaco Glacier, Leifdefjord, Svalbard, Norway 2010

Edwin Giesbers / Orang utan (Pongo pygmaeus) baby sleeping in the arms of an adult, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered

Edwin Giesbers / Orang utan (Pongo pygmaeus) portrait, silhouetted against sky on tree branch, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered

Laurent Geslin / Honey bees (Apis mellifera) flying through the city. Paris, France

Laurent Geslin / Two Kestrels (Falco tinnunculus) on a stone wall in Paris, with tourists in the background. France

Andrew Parkinson / Arctic Skua (Stercorarius parasiticus) pair almost colliding in flight, as they pursue an Arctic Tern (Sterna paradisaea). Shetland Islands, Scotland, UK, July. (non-ex)

Andrew Parkinson / Pair of Great skuas (Stercorarius skua) on a misty moorland with person walking close to their territory. Shetland Islands, Scotland, UK, June. (non-ex)

Floris van Breugel / Kokanee Salmon (Oncorhynchus nerka) on Autumn migration up Taylor Creek from Lake Tahoe, California, USA.

Floris van Breugel / Monarch Butterflies (Danaus plexippus) roosting in winter in dense groups, Southern California, USA

Alex Mustard / A group of Florida manatees (Trichechus manatus latirostrus) sleeping in the afternoon at Three Sisters Spring. Manatees sleep on the bottom of this shallow spring, occasionally rising to the surface...