Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...

Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...

Konrad Wothe / Fritillary butterfly (Argynnis sp) with Scarce copper butterflies (Lycaena virgaureae) Queyras, Alps, France. August.

Nick Upton / Slow motion clip of a Meadow brown butterfly (Maniola jurtina) and a Gatekeeper butterfly (Pyronia tithonus) nectaring on Saw-wort flowers (Serratula tinctoria), with a Silver-washed fritillary (Argyn...

Nick Upton / Close-up of a Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower and flying off, Wiltshire, England, UK, July.

Dave Watts / Dark green fritillary (Argynnis aglaja) on pink clover, south west France.

Jussi Murtosaari / Pallas Fritillary (Argynnis laodice) portrait of female head on, southern Finland, August.

Stephen Dalton / Silver washed fritillary {Argynnis paphia} in flight, UK

Wild Wonders of Europe / Maitland / Round headed club rush (Scirpus holoschoenus) with Heath fritillary butterfly (Melitaea athalia) and Niobe fritillary butterfly (Argynnis niobe var eris) Stenje region, Galicica National Park, Macedon...

Thomas Marent / Silver-washed Fritillary (Argynnis paphia) butterfly, Switzerland

Paul Hobson / Silver washed fritillary {Argynnis paphia} chrysalis / pupa, UK.

David Kjaer / Silver washed fritillary {Argynnis paphia} on flower. UK.