Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male yawning, Tangkoko National Park, Sulawesi, Indonesia.

Markus Varesvuo / Common Raven (Corvus corax) huddled down amidst snow storm, Utajarvi Finland, February

Steven Kazlowski / Lynx (Lynx lynx) adult in the snowy foothills of the Takshanuk mountains, South-East Alaska, USA

David Noton / Rain clouds hanging over the Col de Bavella, Bavella Mountains, Corsica, France. June 2011.

Roy Mangersnes / Mountain tops in clouds at sunset over Raudefjorden, Northern Spitsbergen, Svalbard, Norway, September

Roy Mangersnes / Arctic fox (Alopex / Vulpes lagopus) in tundra landscape, near Longyearbyen, Spitsbergen, Svalbard, Norway, August

Roy Mangersnes / Evening sky over mountain top near Raudefjorden, Northern Spitsbergen, Svalbard, Norway, September

Roy Mangersnes / Delta running through autumn colours in Adventsdalen, Spitsbergen, Svalbard, Norway, August

Roy Mangersnes / Person standing on Pulpit Rock, 604 meters above the Lysefjorden covered in mist, Rogaland, Norway

Roy Mangersnes / Mist above Monaco Glacier, North west of Spitsbergen, Svalbard, Norway, August

Gerrit Vyn / Spotted owl (Strix nebulosa) Gifford-Pinchot National Forest, Washington USA

Alex Mustard / Bluestriped grunt (Haemulon sciurus) portrait, West End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.

Carol Walker / Horse in snow storm with shed in background, USA

George Sanker / Pond in forest, Baxter State Park, Maine, USA, October 2007

Edwin Giesbers / Wild cat (Felis silvestris) in snowstorm captive, Germany

Laurie Campbell / Sandstone cliffs, Rackwick Bay, Hoy, Orkney, Scotland, July 2004

Graham Eaton / View towards Arenig Fach over open moorland, with stormy sky and dark clouds. Snowdonia NP, Gwynedd, Wales, UK, October

Luis Quinta / Abstract image of large sea swells hitting the rocks, Portuguese southwestern coast, Portugal, March, 2010.

Chadden Hunter / King penguin (Aptenodytes patagonicus) chick creche enduring harsh weather, South Georgia. Taken on location for BBC Frozen Planet series, 2008

Loic Poidevin / Redpoll (Carduelis flammea) perched on a branch in snow. Breton Marsh, French Atlantic Coast, March.

Martin Gabriel / Mountain ridges of Caldera de Taburiente silhouetted through clouds. View from Roque de los Muchachos, La Palma Island, Canary Islands, Spain, February.

Sven Zacek / Roe Deer (Capreolus capreolus) herd travelling through snow and fog. Virumaa, Estonia, Europe, March.

Sven Zacek / Heavy rainfall on the calm surface of the Baltic Sea. Hiiumaa Island, Estonia, Europe, August, 2011.

Sven Zacek / Waves crashing onto beach rock under dark clouds. Hiiumaa Island, Estonia, Europe, August 2011.

Nick Garbutt / Emergent Menggaris Tree / Tualang (Koompassia excelsa) protruding from mist and low cloud hanging over lowland rainforest. Danum Valley, Sabah, Borneo.

Ole Jorgen Liodden / Plains Zebra (Equus quagga / quagga) in heavy rain. Masai Mara, Kenya, September.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) colony in heavy snow. South Georgia Island, Southern Ocean, November.

Ole Jorgen Liodden / Brunnich's Guillemot (Uria lomvia) flying between misty colony cliffs. Spitsbergen, Svalbard, August.

Ole Jorgen Liodden / King Penguins (Aptenodytes patagonicus) at sunrise on the coast of South Georgia Island. Southern Ocean, November.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) walking back to colony on South Georgia Island. Southern Ocean, Antarctic Convergence, November.

Ole Jorgen Liodden / Thunderstorm and bushfire at night. Kalahari, Botswana, November.

Ole Jorgen Liodden / Thunderstorm and bushfire at night. Kalahari, Botswana, November.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) emerging from the sea to get back to their colony. South Georgia Island, Southern Ocean, November.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) parents calling and performing 'statue' display at colony. South Georgia Island, Southern Ocean, November.

Peter Lewis / Corfe Castle rising out of mist, viewed from Kingstone, Purbeck, Dorset, UK, September 2010

Fabrice Cahez / Red fox (Vulpes vulpes) pouncing while hunting in field, Vosges, France, August

Steven Kazlowski / Walrus (Odobenus rosmarus) two young bulls, in waters along Spitsbergen and the northwestern coast of the Svalbard Archipelago, Norway, Arctic Ocean, July

Ernie Janes / Three fishing boats wrecked on the coast in winter storms, Pembrokeshire, UK,

Anup Shah / Hippopotamus head portrait (Hippopotamus amphibius) Masai Mara National Reserve, Kenya

Niall Benvie / Beech trees in autumn mist, Morvan regional park, Burgundy, France, November 2010

George Sanker / Limber Pine (Pinus flexilis). Grand Teton National Park, Wyoming, USA, October 2010. Famous patriarch tree of the Grand Tetons just after fall snowstorm. Winner: World Landscapes category in the Outdo...

Grzegorz Lesniewski / Ural Owl (Strix uralensis) perched. Bieszczady National Park, the Carpathians, Poland, December.

Ernie Janes / Storm clouds building up off coasts of Norfolk, Weybourne Cliffs, Norfolk, UK, November

Ernie Janes / Seedling English oak tree (Quercus robur) in the rain, UK

Klaus Echle / Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November.

Radomir Jakubowski / Tree standing in snowy and misty landscape. Rondane National Park, Norway, September 2010. Winner, Fritz Polking Junior Award portfolio, GDT 2011 Competition.

Ernie Janes / Franz Josef Glacier, Southern Alps, South Island, New Zealand

Ernie Janes / Common foxglove (Digitalis purpurea) flowers, UK

David Noton / Rough seas at Durdle Door, Dorset, England, November 2009.

Klaus Echle / Red Fox (Vulpes vulpes) sat on a fallen tree in misty woodlands. Highly Commended in the Animals in their Environment category, Wildlife Photographer of the Year 2011.

Klaus Echle / Badger (Meles meles) foraging on a misty day in woodland clearing, Black Forest, Germany. Highly commended in Animals in their Environment category, Wildlife Photographer of the Year 2011 competition

Philippe Clement / Heavy rain pouring down on lake Lago dei Caprioli surrounded by pine forest. Dolomites, Italy, July 2010.

Philippe Clement / Heavy rain pouring down on lake Lago dei Caprioli surrounded by pine forest. Dolomites, Italy, July 2010.

Sue Daly / The isthmus between Greater and Little Sark, bound in snow. The heaviest snowfall on Sark in living memory. Channel Islands, UK, December 2010.

Sue Daly / John Dory (Zeus Faber) with Pipefish spawn. Channel Islands, UK, August.

Sue Daly / A snow-covered tree by a road. The heaviest snowfall on Sark in living memory. Channel Islands, UK, December 2010.

Luiz Claudio Marigo / A field of Yellow 'Frailejon' (Espeletia grandiflora). Paramo habitat (Andean high altitude fields) Chingaza Natural National Park, municipality of La Calera, Cundinamarca Department, Colombia.

David Woodfall / Sessile Oak (Quercus patraea) in mist. Gilfach Farm SSSI, Radnorshire Wildlife Trust, Wales, UK, November.

Fabrice Cahez / Eurasian Sparrowhawk (Accipiter nisus) rear view perching on fence post with tail feathers spread. Vosges, France, January.

Pete Oxford / Huaorani Indian house. The houses are made by constructing a frame from cut logs and covering it with palm leaves. Gabaro Community, Yasuni National Park, Ecuador, June 2007.

Mark Bowler / Cloud forest Alto Mayo Protection Forest Reserve Amazonas Peru - Habitat of Yellow tailed-woolly monkey. June 2008.

Carol Walker / Andalusian mare running in snow storm, Colorado, USA

Diane McAllister / Brown bear (Ursus arctos) strolling along the beach in the rain, looking towards the camera. Lake Clark National Park, Alaska, USA, July

David Welling / A thunderstorm drops heavy rain over the hoodoo sandstone formations. Bryce Canyon National Park, Utah, USA, August.

Inaki Relanzon / A high view of mist shrouding an autumnal valley. Cadi Natural Park, Catalonia, Barcelona province, Pyrenees, Spain, October 2005.

Inaki Relanzon / Isolated rain storm above the sea. Bahamas Sea, Bahamas, North America, August 2006.

Inaki Relanzon / Dramatic light on Benijo Beach. Anaga Peninsula, Tenerife, Canary Islands, February 2011.

Inaki Relanzon / Limestone formations under a heavy sky. Torcal de Antequera Nature Reserve, Malaga, Spain, November 2009.

Matthew Maran / Dramatic seas crashing onto a rocky shore. The Broken Group Islands from The Wild Pacific Trail, Ucluelet, West coast of Vancouver Island, Canada, September 2010.

David Woodfall / Upland conifer plantation in snow and mist, Gwynedd, North Wales, February 2009

Juergen Freund / The GEO snake expedition team braving the flooded waters to follow the lead of an aboriginal man looking for snakes in their sacred place. Queensland, Australia, February 2008. Model released.

Juergen Freund / Lightning storm over the Australian outback at dusk. Queensland, Australia, February 2008

Anup Shah / Masai giraffe (Giraffa camelopardalis tippelskirchi) crossing the Mara River - wide angle perspective, Masai Mara National Reserve, Kenya. August 2009.

Edwin Giesbers / Mute Swan (Cygnus olor) standing on frosty grass. The Netherlands, March.

Sergey Gorshkov / Salmon (Salmonidae) migrating to spawn in Lake Kuril, Kamchatka, Far East Russia, August

Fergus Gill / Red kite (Milvus milvus) flying in a rain shower. Dumfries and Galloway, Scotland, June

Alex Hyde / Beech woodland in dense fog. Derbyshire, UK, November

Orsolya Haarberg / Reindeer (Rangifer tarandus) walking through snowstorm, Vadso, Varanger, Finnmark, Norway, February 2006

Orsolya Haarberg / Spruce trees covered in hard frost, coniferous forest in winter, Harghita, Romania.

Orsolya Haarberg / Hay meadow, Danube-Drava National Park, Hungary.

Orsolya Haarberg / Icelandic horses out in the snow, Iceland, April 2010

Orsolya Haarberg / Spotted redshank (Tringa erythropus) at shallow water in snow, Ovre-Pasvik National Park, Norway

Orsolya Haarberg / Snow covered landscape with Vik church in the background, the large stones in the foreground have been covered with volcanic ash from the Eyafjallajokull volcano, Iceland, April 2010

Markus Varesvuo / Bearded vulture (Gypaetus barbatus) flying, Spain, November

Anup Shah / Impala herd (Aepyceros melampus) huddled together in the rain, Masai Mara National Reserve, Kenya. April

Charlie Summers / King Penguin (Aptenodytes patagonicus) adult surrounded by huddled chicks, riding out a snowstorm at Gold Harbor, South Georgia.

Andy Rouse / Mountain gorilla (Gorilla beringei) youngster with bad teeth in Susa group, Parc National des Volcans, Rwanda

Angelo Gandolfi / View of mist over mountains, Monte Incudine, Parc Naturel Regional de Corse, Corsica island, France, February 2010

Angelo Gandolfi / Carrion Crows (Corvus corone) mobbing a Bearded vulture (Gypaetus barbatus) Scala di St. Regina gorges. Corte, Parc Naturel Regional de Corse, Corsica island, France, January

Petra Wegner / Two Miniature Schnauzers, black-silver and pepper-and-salt coated, sitting in autumn foliage

Jerry Monkman / Rows of organic Cabbage plants (Brassica oleracea capitata) growing at the Harlow Farm in Westminster, Connecticut River Valley, Vermont, USA. August 2007

Andrew Parkinson / Ptarmigan (Lagopus mutus) in white winter plumage framed by a walker passing by during a blizzard. Cairngorm Mountains, Scotland, UK. February (non-ex)

Andrew Parkinson / Pied wagtail (Motacilla alba yarrellii) standing on snow covered ground, with wind blowing snow around, Derbyshire, UK, January (non-ex)

Andrew Parkinson / Barnacle geese (Branta leucopsis) flock flying in dense winter fog. Dumfries and Galloway, Scotland, UK, January (non-ex)

Andrew Parkinson / Pair of Mallard ducks (Anas platyrhynchos) on a lake in dawn, Derbyshire, UK. April (non-ex)