Guy Edwardes / Dalmatian pelican (Pelecanus crispus), portrait. Lake Kerkini, Sintik, Greece. January.

Jack Dykinga / Greater roadrunner (Geococcyx californianus) running to feed brood with Lizard prey in beak. Catalina State Park, Sonoran Desert, Arizona, USA. June.

George Sanker / Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick looking upwards to parents beak. Atka Bay, Antarctica. August. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

Oscar Dewhurst / Great bittern (Botaurus stellaris) with recently caught fish prey in beak. Suffolk, UK. November.

SCOTLAND: The Big Picture / Grey heron (Ardea cinerea) hunting for food amongst moss with loch and Celtic rainforest in back ground, Scotland, UK, October..

Steven David Miller / Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive.

Marie Read / Ruby-throated Hummingbird (Archilocus colubris), male flying in to feed from cardinal flowers (Lobelia cardinalis) New York, USA. August.

Marie Read / American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April.

Marie Read / California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA

Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA

Marie Read / Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October.