Ismaele Tortella / Eurasian otter (Lutra lutra) hunting in a shoal of Atlantic herring (Clupea harengus), northern Norway, March

Ismaele Tortella / White tailed eagle (Haliaeetus albicilla) catching fish, Lofoten, Norway, August.

Morley Read / Slow motion clip of a Linnaeus' sipo snake (Chironius exoletus) with open mouth, part of threat display, Amazon rainforest, Orellana Province, Ecuador. (non-ex)

Morley Read / Slow motion clip of a large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Morley Read / Parrot snake (Leptophis ahaetulla) threat display, Amazon rainforest, Orellana Province, Ecuador. (non-ex)

Morley Read / Slow motion clip of a group of male Mallards (Anas platyrhynchos) competing violently to mate with a single female, Cornwall, England, UK, May 2018. (non-ex)

Bence Mate / European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition.

Diane McAllister / Polar bears (Ursus maritimus) fighting in water. Beaufort Sea , Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Nature's Best Photography Competition 2019.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) two adults brooding chicks age 5 weeks, Antarctica. Highly commended in the Polar Passion Category of the Nature's Best Photography Awards 2019.

Alex Mustard / Yellowtail tang (Zebrasoma xanthurum) male and female turn belly to belly during spawning to ensure fertilization. Temple, Ras Umm Sid, Sharm El Sheikh, Sinai, Egypt. Red Sea. This action is very fast...

Morley Read / Slow motion clip of a Spiny orb-weaver (Micrathena sp.) weaving a silk nest to protect its eggs, the nest is suspended from a thread to isolate it from predators, Amazon rainforest, Ecuador. Filmed at...

Danny Green / Polar bear (Ursus maritimus) female and cubs feeding on Whale carcass. Svalbard, Norway, July.

Danny Green / Polar bear (Ursus maritimus) and cubs feeding on Whale carcass. Svalbard, Norway, July.

Danny Green / Reindeer (Rangifer tarandus), two play fighting in snow. Svalbard, Norway, April.

Danny Green / Polar bear (Ursus maritimus) female and cubs play fighting. Svalbard, Norway, March 2018.

Danny Green / Polar bear (Ursus maritimus) group on ice, female and cubs feeding on Whale carcass. Svalbard, Norway, June 2018.

Danny Green / Polar bear (Ursus maritimus) hunting Seal on sea ice. Svalbard, Norway, July 2018.

Danny Green / Polar bear (Ursus maritimus), two cubs playing, leaping across sea ice, reflected in water. Svalbard, Norway, July.

Phil Savoie / Blue tit (Cyanistes caeruleus) bringing Crab spider (Misumena vatia) prey to nest, Monmouthshire Wales, UK, May.

Phil Savoie / Smeathman's furrow bee (Lasioglossum smeathmanellum), feeding on wild alpine strawberry (Fragaria vesca), Monmouthshire, Wales, UK. December

Phil Savoie / Scottish wildcat (Felis silvestris grampia) captive, endemic to Scotland. September

Nick Garbutt / King penguin (Aptenodytes patagonicus), two sleeping on snow. Grytviken, South Georgia. March.

John Cancalosi / Sonoran desert toad (Incilius alvarius) pair mating, with trail of spawn in water. Arizona, USA.

Ann & Steve Toon / African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana.

Tony Wu / Humphead parrotfish (Bolbometopon muricatum) shoal spawning in early morning, part of mass aggregation of thousands of fish. Caroline Islands, Palau.

Alex Mustard / Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt.

Alex Mustard / Green turtle (Chelonia mydas) feeding on Seagrass (Halophila stipulacea), blowing sand out of nostrils, on sea floor. Marsa El Shouna, Marsa Alam, Egypt.

Alex Mustard / Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt.

Oscar Dewhurst / Mute swan (Cygnus olor), two cygnets sheltering under parent's wing. Richmond Park, London, England, UK. April.

Clay Bolt / Blue-banded bee (Amegilla sp) males sleeping on stem. Halmahera, North Maluku, Indonesia.

Felis Images / Asiatic lion (Panthera leo persica), female and cubs in morning light, cub biting mother's leg. Gir National Park, Gujarat, India. Photo Phillip Ross/Felis Images

David Tipling / Common European adder (Vipera berus) two males fighting over a female. Holt, North Norfolk, England, UK. April

David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April

David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.

Tony Wu / Humpback whale (Megaptera novaeangliae australis) female with suckling calf. Vava'u Islands, Tonga.

Tony Wu / Sperm whale (Physeter macrocephalus), ten swimming together in gathering of more than 50 individuals. Whales with flatulence and defecating. Sri Lanka.

Jack Dykinga / Greater roadrunner (Geococcyx californianus) running to feed brood with Lizard prey in beak. Catalina State Park, Sonoran Desert, Arizona, USA. June.

George Sanker / Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.

George Sanker / Great egret (Ardea alba) diving head into water to catch prey, in evening light. Myakka River State Park, Florida, USA, March. Sequence 1/2.

Tony Heald / Elephant (Loxodonta africana) babies playing, Etosha National Park,Namibia.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) egg hatching on father's feet. Atka Bay, Antarctica. August. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

Staffan Widstrand / African bush elephant, (Loxodonta africana) two head to head playing, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa

Staffan Widstrand / RF - Pied kingfisher (Ceryle rudis) group of three perched on branch, squabbling, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa

Staffan Widstrand / African Spoonbill (Platalea alba) preening, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa

Sven Zacek / Ural owl (Strix uralensis) in flight, hunting, Tartumaa county, Southern Estonia. February.

Jordi Chias / Chinstrap penguin (Pygoscelis antarcticus) carcass after leopard seal attack, Antarctic Peninsula, Antarctica.

Jordi Chias / Humpback whale (Megaptera novaeangliae) spyhopping, Antarctic Peninsula, Antarctica.

Jordi Chias / Leopard seal (Hydrurga leptonyx) with mouth open, Antarctic Peninsula, Antarctica.

Jordi Chias / Gentoo penguin (Pygoscelis papua) feeding chick, Antarctic Peninsula, Antarctica.

Jordi Chias / Gentoo penguin (Pygoscelis papua) feeding chick, Antarctic Peninsula, Antarctica.

Jordi Chias / Leopard seal (Hydrurga leptonyx) predating a Gentoo penguin (Pygoscelis papua) Antarctic Peninsula, Antarctica.

Jordi Chias / Humpback whale (Megaptera novaeangliae) spyhoping, Antarctic Peninsula, Antarctica.

Ingo Arndt / Female Puma (Puma concolor) with two cubs aged around four months, Torres del Paine National Park, Patagonia, Chile, July.

Matthew Maran / Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. June.

Matthew Maran / Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. July.

Matthew Maran / Red fox (Vulpes vulpes) carrying a large dead Brown rat (Rattus norvegicus) in an urban garden, North London, UK. August.

Matthew Maran / Red fox (Vulpes vulpes) cub defending its kill,a large Brown rat (Rattus norvegicus) from another cub, North London, UK. August. Highly commended in the Behaviour : Mammals Category of the Wildlife Ph...

Tony Wu / North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching, Icy Strait, Alaska, USA. July.

Alex Mustard / Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island. Misool, Raja Ampat, West Papua, Indones...

Alex Mustard / Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the background sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia. Lembeh S...

Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef. This photo shows the larger...

Alex Mustard / Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...

Alex Mustard / Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Robert Valentic / Rough-scaled death adder (Acanthophis rugosus) waving tip of tail as a lure to entice prey within striking range. Mount Isa, Queensland Australia.

Tony Wu / Blacktip reef shark (Carcharhinus melanopterus) beaching itself to catch sardines. Tonga South Pacific.

Tony Wu / View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.

Tony Wu / Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tony Wu / Zoarchias major eelpouts. (Zoarchias major) The orange individual is female, the one in the hole is male. The female has approached the male, curled around him, attempting to dislodge the male from th...

Tony Wu / Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tony Wu / Sardines (Clupeidae) fleeing from predators in the early morning. They were being herded and chased by Trevallies and Blacktip reef sharks in the water, and picked off by brown boobies and terns from...

Lucas Bustamante / Giant river otter (Pteronura brasiliensis) eating a catfish. Yasuni National Park, Orellana, Ecuador

Lucas Bustamante / Treehopper (Alchisme onorei) with eggs Papallacta, Napo, Ecuador

Lucas Bustamante / Marine Iguanas (Amblyrhynchus cristatus) basking. Fernandina Island, Galapagos Islands.

Nick Upton / Juvenile Goldenrod spider (Misumena vatia) camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, reacting to and catching a fly (Muscidae), Wiltshire, England, UK, June.

Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.

David Gallan / Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement. Highly Commended 2018 Wildlife Photographer Of The Yea...

Nick Upton / Forest caterpillar hunter (Calosoma sycophanta) feeding on a Gypsy moth caterpillar (Lymantria dispar) it has killed in a Holm oak tree, Bacu Goloritze ravine, Baunei, Sardinia, Italy, May.

John Chan / Crenulated fiddler crab (Uca crenulata) waving its major claw at another crab, part of a courtship display, Southern California, USA, June.

John Chan / Peregrine falcon (Falco peregrinus) plucking Dowitcher (Limnodromus) prey, Southern California, USA.

John Chan / Killdeer (Charadrius vociferus) stamping its feet to create vibrations in the sand as it forages for insects, Southern California, USA, August.

David Woodfall / Little tern (Sterna albifrons ) feeding sand eel (Hyperoplus spp) to young chick. Gronant Dunes, Denbighshire, Wales, UK, June.

John Shaw / Common eider duck (Somateria mollissima) female defending her nest from arctic fox (Vulpes lagopus). Longyearbyen, Spitsbergen. June.

John Shaw / Great frigatebird (Fregata minor) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May.

John Shaw / Osprey (Pandion haliaetus) with Atlantic needlefish (Strongylura marina) it has caught. Blue Cypress Lake, Florida, USA, April.

John Shaw / Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November.

John Shaw / Namaqua chameleon (Chamaeleo namaquensis) catching larvae with its extended tongue. In the Namib desert near Swakopmund, Namibia.

Andy Rouse / Barn owl (Tyto alba) hunting with sunset light on wings, UK. March.

Shane Gross / Elkhorn corals (Acropora palmata) at night silhouetted by a scientist's light during the annual coral spawn. Abaco, Bahamas.

Espen Bergersen / Humpback whales (Megaptera novaeangliae) showing tail fluke when they dive. Kvanangen, Troms, Norway. December

Espen Bergersen / Humpback whale (Megaptera novaeangliae) feeding around herring fishing vessel. Kvanangen, Troms, Norway. November

Espen Bergersen / Humpback whale (Megaptera novaeangliae) diving. In background a diving boat and 3 divers visible in the water. Kvanangen, Troms, Norway. November

Espen Bergersen / Aerial view of Fin whale (Balaenoptera physalus). Kvanangen, Troms, Norway. November