Ismaele Tortella / Northern gannet (Morus bassanus) colony, chick climbing back into nest, with the midnight sun, Norway, July

Heike Odermatt / Brunnich's Guillemot (Uria lomvia) nesting colony on cliff, Alkefjellet, Hinlopen Strait, Svalbard, Norway

Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled together under the Southern Lights, Aurora australis, Atka Bay, Antarctica. September.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

John Shaw / Roseate spoonbill (Platalea ajaja) at its nesting colony site, in early morning light. St. Johns Management Area, Florida, USA, April.

Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony with chicks, Atka Bay, Queen Maud Land, Antarctica. October.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony with with creche of huddling chicks, Atka Bay, Queen Maud Land, Antarctica. October.

Nick Upton / White stork (Ciconia ciconia) parent standing beside its three developing chicks. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cotswold...

Nick Upton / White stork (Ciconia ciconia) nest at captive breeding colony raising chicks for UK White Stork reintroduction project at the Knepp Estate. Cotswold Wildlife Park, Oxfordshire, UK, May 2019.

Nick Upton / Richard Wardle feeding fish to recently hatched White stork (Ciconia ciconia) chicks in nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Esta...