Stefan Christmann / Emperor penguin (Aptenodytes forsteri) footprints on sea ice. Atka Bay, Antarctica. August.
01633174
Colin Varndell / Song thrush (Turdus philomelos) anvil with broken snail shells. Dorset, England, UK. April.
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Sven Zacek / Ural owl (Strix uralensis) print in snow after landing on prey, Valgamaa county, Southern Estonia. January
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) tracks in snow, Atka Bay, Queen Maud Land, Antarctica. October.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) tracks in snow, Atka Bay, Queen Maud Land, Antarctica. October.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) footprints in the snow. Atka Bay, Antarctica. July.
01619878
Stefan Christmann / Sea ice with hollows where Emperor penguin (Aptenodytes forsteri) breeding colony stood whilst huddling. Atka Bay, Antarctica. July.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) footprints in snow. Atka Bay, Antarctica. August.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) footprints on sea ice. Atka Bay, Antarctica. August.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) tracks in snow, during polar night. Atka Bay, Antarctica. June.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) foorprints in snow, during polar night. Atka Bay, Antarctica. June.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) foorprints in snow, during polar night. Atka Bay, Antarctica. June.
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Karine Aigner / Indian peafowl (Pavo cristatus), peacock walking across track between flowering Bengal kino / Flame of the forest (Butea monosperma) trees. Ranthambore National Park, Rajasthan, India.
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Sue Flood / Emperor penguins (Aptenodytes forsteri) tracks from tobogganing, Antarctica.
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Nick Hawkins / Piping plover (Charadrius melodus) tracks in sand, Nova Scotia, Canada, May.
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Lynn M. Stone / Dunes and vegetative cover along Tampa Bay with tracks of Yellow-crowned night heron (Nyctanassa violacea) a predator of Ghost Crabs (Ocypodinae) that live in dune burrows), Muillet Key, St. Petersbur...
01571992
Enrique Lopez-Tapia / Wading bird footprints in sand on beach, Orango Islands National Park, Bijagos UNESCO Biosphere Reserve, Guinea Bissau.
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Enrique Lopez-Tapia / Wading bird footprints in sand on beach, Orango Islands National Park, Bijagos UNESCO Biosphere Reserve, Guinea Bissau.
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Enrique Lopez-Tapia / Wading bird footprints in sand on beach, Orango Islands National Park, Bijagos UNESCO Biosphere Reserve, Guinea Bissau.
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Tui De Roy / Erect-crested penguins (Eudyptes sclateri) feathers left behind after seasonal moulting. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered specie...
01499974
Tui De Roy / Erect-crested penguins (Eudyptes sclateri) feathers left behind after seasonal moulting. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered specie...
01498033
Roger Powell / Female Bearded Tit (Panurus biarmicus) feeding on grit along a track by a reed bed. Leighton Moss, Lancashire, UK. October.
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SCOTLAND: The Big Picture / Ptarmigan (Lagopus mutus) tracks running up through a boulder field, Cairngorms National Park, Scotland, UK, March 2017.
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Jean E. Roche / Spruce (Picea abies) wood on ground after Black woodpecker (Dryocopus martius) has been drilling trunk for prey. Jura Mountains, Switzerland, March.
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Jean E. Roche / Spruce (Picea abies) wood on ground after Black woodpecker (Dryocopus martius) has been drilling trunk for prey. Jura Mountains, Switzerland, March
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Sam Hobson / Seabird researchers track GPS logged Northern gannets (Morus bassanus). Ailsa Craig, Scotland, UK. August 2015.
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Sam Hobson / Seabird researchers track GPS logged Northern gannets (Morus bassanus). Ailsa Craig, Scotland, UK. August
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Richard Du Toit / Habitat burnt after wildfire, Marievale Bird Sanctuary, South Africa, June.
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Juergen Freund / Southern Cassowary (Casuarius casuarius) claw print next to human hand. Queensland, Australia.
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David Tipling / RF - King penguin (Aptenodytes patagonicus) foot prints in sand, St Andrew's Bay, South Georgia. January. (This image may be licensed either as rights managed or royalty free.)
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Pete Oxford / Red junglefowl (Gallus gallus) on track, Kaziranga National Park, Assam, North East India.
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Michael Quinton / Canada Goose (Branta canadensis) tracks in snow, Alaska
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Markus Varesvuo / Black grouse (Tetrao / Lyrurus tetrix) foot prints on snow, Utajarvi, Finland, April.
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Markus Varesvuo / Capercaillie (Tetrao urogallus) tracks of a displaying male, Kuusamo, Finland, March.
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Markus Varesvuo / Willow grouse / Ptarmigan (Lagopus lagopus) feeding tracks under a Birch tree (Betula sp), Inari Kiilopaa, Finland, January.
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Brent Stephenson / Researcher using radio-telemetry equipment to track a Northern brown kiwi (Apteryx mantelli) fitted with a radio transmitter Beside her stands a trained kiwi-tracking dog, Cape Kidnappers, Hawkes Bay,...
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Donald M. Jones / Wild Turkey (Meleagris gallopavo) tracks in snow, western Montana
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Bruno D'Amicis / Human footprints and wading bird footprints in cracked mud, Shesher lake, Fogera Plains, Lake Tana Biosphere Reserve, Ethiopia. December 2013.
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Chien Lee / Satanic Nightjar (Eurostopodus diabolicus) in defensive posture, Anaso Track, Sulawesi, Indonesia
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Chien Lee / Satanic Nightjar (Eurostopodus diabolicus) camouflaged in leaf litter, Anaso Track, Sulawesi, Indonesia
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Dong Lei / Lady Amhersts pheasant (Chrysolophus amherstiae) female walking on track with Buddhist prayer flags, Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China. May.
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Donald M. Jones / Canada Goose (Branta canadensis) tracks in snow, North America
00546207
Stefan Christmann / Emperor Penguin (Aptenodytes forsteri) tracks in snow, Queen Maud Land, Antarctica
00523171
Kevin Schafer / Black-footed Penguin (Spheniscus demersus) footprints in sand, Boulders Beach, Cape Peninsula, South Africa
00476276
Martin Willis / Banded Whiteface (Aphelocephala nigricincta) male perched in shrub, Strzelecki track, South Australia, Australia
01291286
Steven David Miller / Australasian Grebe (Tachybaptus novaehollandiae) Non-breeding or immature bird on borewater wetland, Muloorina Station, Oodnadatta Track, South Australia
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Sergey Gorshkov / Raptor imprint in snow, left after bird foraged for prey hidden beneath the snow, Kamchatka, Russia
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Colin Monteath / New Zealand Robin (Petroica australis), Inland Pack Track, Paparoa National Park, New Zealand
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Richard Du Toit / Road through savanna grass, Marievale Bird Sanctuary, South Africa
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Richard Du Toit / Road through savanna grass, Marievale Bird Sanctuary, South Africa
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Jasper Doest / Sanderling (Calidris alba) tracks in the mud, Noord-Holland, Netherlands
00429963
Konrad Wothe / Eurasian Kestrel (Falco tinnunculus) print left in snow after catching a mouse, Germany
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Sebastian Kennerknecht / Snowy Plover (Charadrius nivosus) breeding sign with footsteps, Scott Creek Beach, Santa Cruz, California
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Donald M. Jones / Wild Turkey (Meleagris gallopavo) tracks in snow, western Montana
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Jasper Doest / Northern Hawk Owl (Surnia ulula) imprint in the snow, Norway
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Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) walking on steep and snowy alpine slope, Antarctic Bay, South Georgia Island
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Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) walking through snow, Antarctic Bay, South Georgia Island
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Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) trio on beach, St. Andrews Bay, South Georgia Island
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Jan Vermeer / Ring-necked Pheasant (Phasianus colchicus) footprints in the sand, Europe
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Gerry Ellis / Bird tracks in sand dune, Coral Pink Sand Dunes State Park, southern Utah
00161018
Michael Quinton / Western Sandpiper (Calidris mauri) footprints and holes from probing beaks on mudflats during spring migration stop-over at the Copper River Delta, Alaska
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Jim Brandenburg / Common Raven (Corvus corax) tracks in sand, Monument Valley National Monument, Arizona
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Tui De Roy / Rockhopper Penguin (Eudyptes chrysocome) group running on beach, Saunders Island, Falkland Islands
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Tui De Roy / Rockhopper Penguin (Eudyptes chrysocome) group running on beach, Saunders Island, Falkland Islands
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Agustin Esmoris / Lesser Rhea (Rhea pennata) footprint, Argentina
00544515
Tui De Roy / Okarito Kiwi (Apteryx rowi) radio transmitters used to track individuals, West Coast Wildlife Centre, New Zealand
00512725
D. Parer & E. Parer-Cook / Superb Lyrebird (Menura novaehollandiae) male crossing bush track, Sherbrooke Forest Park, Victoria, Australia
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D. Parer & E. Parer-Cook / Little Blue Penguin (Eudyptula minor) walking track worn in by trips from nests inland, Phillip Island, Australia
00455662
Kevin Schafer / Southern Cassowary (Casuarius casuarius) footprints on sandy beach, Atherton Tableland, Queensland, Australia
00420016
Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) walking on beach leaving footprints in sand, St. Andrews Bay, South Georgia Island
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Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) tracks in fresh spring snow, Salisbury Plain, South Georgia Island
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Jim Brandenburg / Grey Heron (Ardea cinerea) pair walking across desert sand dune, Namib Desert, Namibia
00141320
Tui De Roy / Laysan Albatross (Phoebastria immutabilis) tracks on beach around Scaevola bushes, Midway Atoll, Hawaii
00076750
Jim Brandenburg / Bird tracks in red sand, Texas
00114262
Mitsuaki Iwago / Adelie Penguin (Pygoscelis adeliae) in snow, Antarctica
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Roger Powell / Willow grouse (Lagopus lagopus) male in winter plumage, walking across a snowy forest track, Pokka, Lapland, Finland. May.
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Russell Laman / Adelie penguin (Pygoscelis adeliae) walking over snow leaving footprints, Cape Evans, Ross Island, McMurdo Sound, Ross Sea, Antarctica.
01728656
Ripan Biswas / A bag confiscated from a poacher containing dead birds including Red-headed swamp hen (Porphyrio sp.), Bronze-winged jacana (Metopidius indicus), Cotton pygmy goose (Nettapus coromandelianus), Eurasia...
01716785
Tui De Roy / Galapagos penguin (Spheniscus mendiculus) diving after small fish into shallow waters. Galapagos Islands, Ecuador. Pacific Ocean.
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Espen Bergersen / Four Rock ptarmigans (Lagopus muta) resting on snow-covered mountainside at dusk, close to Tromso city, Norway. March.
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Espen Bergersen / Aerial view of a dry lake with colourful ponds surrounded by bird tracks, Hungary. February, 2019.
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Tui De Roy / Aerial view of female Olive ridley turtle (Lepidochelys olivacea) departing beach after nesting during massive arribada, with Black vultures (Coragyps atratus) looking out for opportunities to scaveng...
01716894
Tui De Roy / Aerial view of massive arribada of Olive ridley turtle (Lepidochelys olivacea), with over 300,000 females coming ashore to nest on 3 km of 15 km beach over period of three days and nights, with flock...
01716590
Tui De Roy / Blue-footed booby (Sula nebouxii) diving after small shoal of fish along deep sea wall. Galapagos Islands, Ecuador.
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Roland Seitre / Houbara bustard (Chlamydotis undulata) tracks across sandy desert floor, Algeria.
01717857
Roland Seitre / Red-legged seriema (Cariama cristata) running along sandy track, Brazil.
01716507
Roland Seitre / Siberian crane (Grus leucogeranus) standing on one leg in snow, Russia. Captive. Critically endangered.
01716332
Roland Seitre / Madagascar partridge (Margaroperdix madagarensis) pair, male in foreground, standing on sandy track, Madagascar. Captive.
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Markus Varesvuo / Male Rock ptarmigan (Lagopus mutus) in reindeer hoof print, Utsjoki Finland, February. Highly Commended in the Behaviour: Birds category of the Wildlife Photographer of the Year Competition 2022. When...
01687262
Roland Seitre / Lone Adelie penguin (Pygoscelis adeliae) and its trail in the snow appearing lost, 10km from sea, Antarctica.
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Roland Seitre / Lone Adelie penguin (Pygoscelis adeliae) tobogganing on ice shelf, Antarctica.