Stefan Christmann / Emperor penguin (Aptenodytes forsteri), four journeying across sea ice to feed young. Atka Bay, Antarctica. August.

Tui De Roy / Emperor penguin (Aptenodytes forsteri), Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November.

Tui De Roy / Emperor penguin (Aptenodytes forsteri) chicks, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November.

Paul Harcourt Davies / Newly emerged adult Two-tailed Pasha (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August

Juergen Freund / Detail of Emperor angelfish (Pomacanthus imperator) Bali, Indonesia, Indo-pacific

Stephen Dalton / Emperor dragonfly (Anax imperator) in flight, UK

Doug Allan / Emperor penguins (Aptenodytes forsteri) swimming underwater, Cape Washington, Antarctic, Ross Sea, October

Stephen Dalton / Emperor dragonfly {Anax imperator} male in flight, UK

Doug Allan / Emperor penguins {Aptenodytes forsteri} diving, Cape Washington, Antarctica. Freeze Frame book plate page 133.

Ross Hoddinott / Female emperor dragonfly (Anax imperator) laying eggs at the edge of a pond. Cornwall, UK.

Ross Hoddinott / Female emperor dragonfly (Anax imperator) laying eggs at the edge of a pond. Cornwall, UK.

Hans Christoph Kappel / Lesser Purple Emperor butterfly.