Ross Hoddinott / Red fox (Vulpes vulpes) vixen yawning in the morning light (captive). Lifton, Devon, UK, May

Marie Read / Tufted puffin (Fratercula cirrhata), close-up portrait, St. Paul Island, Pribilofs, Alaska, USA, July.

Solvin Zankl / Frilled shark (Chlamydoselachus anguineus) specimen with mouth open, from Atlantic Ocean, at a depth of 729m.

Pete Oxford / Galapagos sealion (Zalophus wollebaeki) mother and pup resting on beach, Galapagos.

Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month cubs with mother 'T19 Krishna' in background, Ranthambhore National Park, India.

Fiona Rogers / Tana mangabey (Cercocebus galeritus) adult and juvenile sitting - portrait. Tana River Forest, South eastern Kenya.

Ben Cranke / Walrus (Odobenus rosmarus) portrait, Svalbard, Norway. July.

Anup Shah,Anup Shah / Lioness (Panthera leo) resting on a fallen tree with a courting male. Masai Mara National Reserve, Kenya, July

Konrad Wothe / Common green tree frog (Litoria caerulea) in rainforest, Iron Range National Park, Cape York Peninsula, North Queensland, Australia, October

Klaus Echle / Lynx (Lynx lynx) cub calling. Hannau, Germany, September. Captive

Ernie Janes / Black labrador dog resting on reeds used by thatcher to thatch roof, Norfolk, UK, November

Ernie Janes / Basset hound, portrait, resting head on back of car, eyes half closed, UK