Michael Pitts / Gentoo penguins (Pygoscelis papua) calling and defending their eggs from Southern Skua (Stercorarius antarcticus) hunting for unattended eggs or chicks, Sea Lion island, The Falklands, South Atlantic.

David Tipling / Great Skua (Stercorarius skua) in flight low over water, Fetlar, Shetland, June

Ole Jorgen Liodden / Brown Skua (Stercorarius antarcticus) bathing, Auckland Islands. New Zealand. November.

Ole Jorgen Liodden / Brown Skua (Stercorarius antarcticus) bathing, Auckland Islands. New Zealand. November.

Andy Rouse / Lonnberg's / Subantarctic Skua (Stercoraius antarcticus lonnbergi) displaying aggressive behaviour, South Shetland Islands, Antarctica. (Non-ex)

Andrew Parkinson / Great Skua (Stercorarius skua) territorial adult displaying to other Skuas passing overhead by stretching out its wings and calling. Shetland Islands, Scotland, UK, June. (non-ex)

David Tipling / Falkland / Brown skua (Stercorarius antarcticus) in flight carrying a stolen Gentoo penguin egg (Pygoscelis papua) Sea Lion Island, Falklands

Hanne & Jens Eriksen / Antarctic skua aggressive display. South Georgia {Stercorarius antarcticus}

Jessie Lane / Antarctic skua {Stercorarius antarcticus} South Georgia (Godthul harbour in background)

Ashley Cooper / Brown skua (Stercorarius antarcticus) on the beach on Prion Island, South Georgia, Southern Ocean.

Tony Heald / Brown skua (Stercorarius antarcticus) resting on tussock grass. Right Whale Bay, South Georgia. September.

Tony Heald / Brown skua (Stercorarius antarcticus), two fighting over carrion whilst standing in sea. South Georgia. September.