John Shaw / Sally lightfoot crabs (Grapsus grapsus) on the beach at Puerto Egas, Santiago Island, Galapagos, May.

Pascal Kobeh / Convex reef crab (Carpilius convexus) at night, Sulu Sea, Philippines

Michele Westmorland / Batwing coral crab (Carpilius corallinus) on coral reef, St Vincent and the Grenadines, Caribbean, May

Luciano Candisani / Batwing Coral Crab (Carpilius corallinus) face, Rocas Atoll, Brazil

Gary Bell/ Oceanwide / Seven-eleven Crab (Carpilius maculatus), Great Barrier Reef, Australia

Pascal Kobeh / A Batwing or Queen crab (Carpilius corallinus) among corals, The Gardens of the Queen, Cuba, Caribbean Sea.

Pascal Kobeh / Convex reef crab (Carpilius convexus) two fighting for territory at night. Philippines.