Phil Savoie / Blue tit (Cyanistes caeruleus) bringing Crab spider (Misumena vatia) prey to nest, Monmouthshire Wales, UK, May.

Phil Savoie / Yellow legged mining bee (Andrena flavipes) female at nest burrow, River Monnow, Monmouthshire, Wales, UK, April.

Phil Savoie / Painted nomad bee (Nomada fucata) male bee with characteristic green eyes, sleeping with jaws locked around grass stem, Monmouthshire, Wales, UK. June

Daniel Heuclin / Llama (Lama glama) Laguna colorada. Altiplano, Bolivia.

Daniel Heuclin / Llama (Lama glama) herd, Laguna colorada. Altiplano, Bolivia.

Phil Savoie / Grey patched mining bee (Andrena nitida),male feeding rambling rose flower (Rosa sp) Monmouthshire, Wales, UK. April

Phil Savoie / Grey-patched Mining Bee (Andrena nitida) feeding on Dandelion (Taraxacum offinicale) Monmouthshire, Wales, UK. April

Matthew Maran / Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. June.

Matthew Maran / Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. July.

Matthew Maran / Red fox (Vulpes vulpes) carrying a large dead Brown rat (Rattus norvegicus) in an urban garden, North London, UK. August.

Matthew Maran / Red fox (Vulpes vulpes) cub defending its kill,a large Brown rat (Rattus norvegicus) from another cub, North London, UK. August. Highly commended in the Behaviour : Mammals Category of the Wildlife Ph...

Tony Wu / Humpback whale (Megaptera novaeangliae) lying on its side while executing a series of pectoral fin slaps on ocean surface. Vava'u, Tonga, South Pacific.

Tony Wu / Humpback whale (Megaptera novaeangliae australis) female breaching. Vava'u, Tonga. Pacific Ocean.

Tony Wu / Humpback whale calf (Megaptera novaeangliae australis) female nudging her mother's mammary gland whilst trying to feed. Vava;u, Tonga, South Pacific.

Tony Wu / North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching, Icy Strait, Alaska, USA. July.

Alex Mustard / Split level image of Leather corals (Sarcophyton sp.) flourishing in shallow water close to uninhabited coral islands. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean...

Alex Mustard / Commensal emperor shrimp (Periclimenes imperator) moves across the colourful red patterned surface of its host Candycane sea cucumber (Thelenota rubralineata). Misool, Raja Ampat, West Papua, Indonesi...

Alex Mustard / Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island. Misool, Raja Ampat, West Papua, Indones...

Alex Mustard / California sea lion (Zalophus californianus) pups in an underwater cave. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.

Alex Mustard / Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind , in the plankton-rich green waters of La Paz bay, Baja...

Alex Mustard / Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean

Alex Mustard / School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean

Alex Mustard / Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the background sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia. Lembeh S...

Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Thorny seahorse (Hippocampus histrix) gripping onto a sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef. This photo shows the larger...

Alex Mustard / Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...

Alex Mustard / Common seahorse (Hippocampus kuda) female wrapping her prehensile tail around piece of wood on the seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Veined octopus (Amphioctopus marginatus) shelters inside an old clam shell, with silver-sides in the water above. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Veined octopus (Amphioctopus marginatus) emerging from its home in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Staffan Widstrand / Grey francolin (Francolinus pondicerianus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Indian peafowl or blue peafowl (Pavo cristatus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India

Staffan Widstrand / Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India

Robert Valentic / Central bearded dragon (Pogona vitticeps) basking in mallee / heathland habitat on the Murrayville Track in the Big Desert Wilderness of north-western Victoria, Australia.

Robert Valentic / Heath Monitor (Varanus rosenbergi) male enacting a threat display, in mallee/heath habitat near Pinnaroo, South Australia.

Robert Valentic / Rough-scaled death adder (Acanthophis rugosus) waving tip of tail as a lure to entice prey within striking range. Mount Isa, Queensland Australia.

Robert Valentic / Striped legless lizard (Delma impar) female, in volcanic plain near Reedesdale in central Victoria, Australia.

Loic Poidevin / Northern ravens (Corvus corax) interacting in snow, Leon, Spain, February.

Loic Poidevin / Eurasian blue tit (Cyanistes caeruleus) on a branch, Vendee, France, January.

Tony Wu / Fringed blenny (Chirolophis japonicus) guarding its eggs. Miyagi Prefecture, Japan.

Tony Wu / Blacktip reef shark (Carcharhinus melanopterus) beaching itself to catch sardines. Tonga South Pacific.

Tony Wu / View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.

Tony Wu / Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tony Wu / Zoarchias major eelpouts. (Zoarchias major) The orange individual is female, the one in the hole is male. The female has approached the male, curled around him, attempting to dislodge the male from th...

Tony Wu / Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tony Wu / Sardines (Clupeidae) fleeing from predators in the early morning. They were being herded and chased by Trevallies and Blacktip reef sharks in the water, and picked off by brown boobies and terns from...

Alex Mustard / Shoal of three Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Portrait of a Giant frogfish (Antennarius commersoni) on a large Yellow elephant ear sponge (Ianthella basta). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Portrait of a tarpon (Megalops atlanticus) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.

Alex Mustard / Blue whale (Balaenoptera musculus) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka.

Alex Mustard / Blue whale (Balaenoptera musculus) swims beneath the surface of the ocean. Indian Ocean, off Sri Lanka.

Alex Mustard / Pod of False killer whales (Pseudorca crassidens) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka.

Alex Mustard / Forster's hawkfish (Paracirrhites forsteri) on a coral reef. Jackson Reef, Tiran, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Egypt.

Alex Mustard / Coral (Lobophytum sp.) on a reef flat, with Hawksbill turtle (Eretmochelys imbricata) in background. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Alex Mustard / Hawksbill turtle (Eretmochelys imbricata) unusually feeding on hard coral polyps. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Alex Mustard / Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Alex Mustard / Carapace of a dead olive Ridley turtle (Lepidochelys olivacea) entangled in discarded fishing gear. Indian Ocean, off Sri Lanka.

Alex Mustard / Green turtle (Chelonia mydas) descending after breathing at the surface. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Lucas Bustamante / Labiated rainfrog (Pristimantis labiosus) feeding on spider prey, Mashpi, Pichincha, Ecuador

Lucas Bustamante / Giant river otter (Pteronura brasiliensis) eating a catfish. Yasuni National Park, Orellana, Ecuador

Lucas Bustamante / Toque Macaques (Macaca sinica) Polonnaruwa, Central Province, Sri Lanka

Lucas Bustamante / Anhinga (Anhinga anhinga). Yasuni National Park, Orellana, Ecuador

Lucas Bustamante / Golden-Tailed Sapphire (Chrysuronia oenone) Sumaco National Park, Napo, Ecuador

Lucas Bustamante / Lesser Clownfrogs (Dendropsophus parviceps) with their eggs. Sumaco National Park, Napo, Ecuador

Lucas Bustamante / Lichen Katydid (Markia hystrix) camouflage in lichens. Sumaco National Park, Napo, Ecuador

Lucas Bustamante / Tropical shield mantis (Choeradodis sp.) Sumaco National Park, Napo, Ecuador

Lucas Bustamante / Treehopper (Alchisme onorei) with eggs Papallacta, Napo, Ecuador

Lucas Bustamante / Green-crowned woodnymph (Thalaurania fannyi) Mashpi, Pichincha, Ecuador

Lucas Bustamante / Choco Toucan (Ramphastos brevis) on a tree with epiphytes, Mashpi, Pichincha, Ecuador

Lucas Bustamante / Sword-billed hummingbird (Ensifera ensifera) Yanacocha, Pichincha, Ecuador

Lucas Bustamante / Marine Iguanas (Amblyrhynchus cristatus) basking. Fernandina Island, Galapagos Islands.

Lucas Bustamante / Sri Lankan leopard (Panthera pardus kotiya) rolling on its back, Yala National Park, Sri Lanka

Lucas Bustamante / Sri Lankan elephant (Elephas maximus maximus) walking past tourist jeep with tourists, Yala National Park, Southern Province, Sri Lanka

Lucas Bustamante / Jaguar (Panthera onca) standing in mud, Yasuni National Park, Orellana, Ecuador

Lucas Bustamante / Zigzag Heron (Zebrilus undulatus) camouflaged in branches while nesting. Yasuni National Park, Ecuador

Lucas Bustamante / Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador

Fabio Pupin / Onager (Equus hemionus), Negev desert, Israel, April.

Fabio Pupin / Red swamp crayfish (Procambarus clarcki), invasive species, captive, Italy, July.

Eladio Fernandez / Fernandina's flicker (Colaptes fernandinae) at hole in tree, Cuba. Vulnerable species.

Eladio Fernandez / Blue-headed quail-dove (Starnoenas cyanocephala) profile, Cuba. Endangered species. Endemic.

David Gallan / Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement. Highly Commended 2018 Wildlife Photographer Of The Yea...

Franco Banfi / Sleeping sperm whale, (Physeter macrocephalus) Dominica, Caribbean Sea, Atlantic Ocean.

Franco Banfi / Sperm whale, (Physeter macrocephalus) Dominica, Caribbean Sea, Atlantic Ocean.

Franco Banfi / Pod of sperm whale socializing, (Physeter macrocephalus), Dominica, Caribbean Sea, Atlantic Ocean.

Oriol Alamany / Snow leopard (Panthera uncia) female with juvenile, walking in snow, in Spiti Valley, Cold Desert Biosphere Reserve, Himalaya, Himachal Pradesh, India, March

Oriol Alamany / Snow leopard (Panthera uncia) male in snow, in Spiti Valley, Cold Desert Biosphere Reserve, Himalaya, Himachal Pradesh, India, March

Oriol Alamany / Snow leopard (Panthera uncia) female (hiding below the rock) with her two juveniles resting in snow, in Spiti Valley, Cold Desert Biosphere Reserve, Himalaya, Himachal Pradesh, India, March

Jack Dykinga / Sandhill cranes (Antigone canadensis) and Snow geese (Chen caerulescens) at sunrise, Bosque del Apache National Wildlife Refuge, New Mexico, USA, November.

Jack Dykinga / Bald eagle (Haliaeetus leucocephalus) perched as Snow geese (Chen caerulescens) fly by, Bosque del Apache National Wildlife Refuge, New Mexico, USA, January.

Jack Dykinga / Snow goose (Chen caerulescens) landing into a tight packed flock on lake, Bosque del Apache National Wildlife Refuge, New Mexico, USA. January.

Jack Dykinga / Sandhill cranes (Antigone canadensis) feeding in pond, Bosque del Apache National Wildlife Refuge, New Mexico, USA. January.

Jack Dykinga / Snow goose (Chen caerulescens) landing into flock, Bosque del Apache National Wildlife Refuge, New Mexico, USA, January.

Jack Dykinga / Sandhill cranes (Antigone canadensis) taking off and flying past a full 'super-moon' en route to feeding in neighboring fields. Bosque del Apache National Wildlife Refuge, New Mexico, USA, Jan...