Stephane Granzotto / Alpine marmot (Marmota marmota) standing alert outside burrow, takes fright and runs inside, Rhone-Alpes, France, July.
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Jussi Murtosaari / Peacock Butterfly (Inachis io) closeup of wings, South Karelia, southern Finland, September.
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Edwin Giesbers / Close-up of an eye of a Sumatran tiger (Panthera tigris sumatrae), captive, native to Sumatra, Indonesia.
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Edwin Giesbers / Close-up portrait of a European grey wolf (Canis lupus), captive, Norway, February.
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Edwin Giesbers / Face portrait of a European lynx (Lynx lynx), captive, Norway, February.
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Radomir Jakubowski / Muskox (Ovibos moschatus) closeup resting, Dovrefjell national park, Norway, September
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Marie Read / American oystercatcher (Haematopus palliatus), closeup of individual feeding by using its chisel-shaped bill to pry open shellfish, Fort De Soto Park, St. Petersburg, Florida, USA, March.
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David Fleetham / Painted frogfish (Antennarius pictus) with first dorsal spine and lure extended to attract prey. Dumaguete, Philippines.
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David Fleetham / Imperial commensal shrimp (Periclimenes imperator / Zenopontonia rex) on Spanish dancer nudibranch (Hexabranchus sanguineus) at night. Philippines.