Ismaele Tortella / Northern gannet (Morus bassanus) colony, chick climbing back into nest, with the midnight sun, Norway, July

Heike Odermatt / Brunnich's Guillemot (Uria lomvia) nesting colony on cliff, Alkefjellet, Hinlopen Strait, Svalbard, Norway

Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled together under the Southern Lights, Aurora australis, Atka Bay, Antarctica. September.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

John Shaw / Roseate spoonbill (Platalea ajaja) at its nesting colony site, in early morning light. St. Johns Management Area, Florida, USA, April.

Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony with chicks, Atka Bay, Queen Maud Land, Antarctica. October.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony with with creche of huddling chicks, Atka Bay, Queen Maud Land, Antarctica. October.

Alex Hyde / Colonies of King penguins (Aptenodytes patagonicus) and Southern Elephant Seals (Mirounga leonina) on beach at St. Andrews Bay, South Georgia. November.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony, males huddling on sea ice during polar night, incubating eggs, ice shelf in background. Atka Bay, Antarctica. July 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled during polar night, males incubating eggs. Atka Bay, Antarctica. July 2017.