Bence Mate / Collared Aracari (Pteroglossus torquatus) displaying at Montezuma Oropendola (Psarocolius montezuma) Laguna del Lagarto, Santa Rita, Costa Rica

Shattil & Rozinski / Sharp tailed grouse {Tympanuchus phasianellus} males fighting on lek, Colorado, USA

Shattil & Rozinski / Gunnison sage grouse {Centrocercus minimus} males displaying at lek, USA, Endangered

David Tipling / Black Cock / Grouse {Tetrao tetrix} fighting on lek in snow, Finland. Males competiong to mate with females

John Cancalosi / Brown vine snake, defensive display {Oxybelis aeneus} Costa Rica

DAVID TIPLING / Red-throated diver {Gavia stellata} threat posture, Finland.

Richard Du Toit / Two male Hippopotamus (Hippopotamus amphibius) sparring, Moremi reserve, Botswana, Southern Africa

Miles Barton / Sichuan golden monkey male yawn threat display (Rhinopithecus roxellana roxellana) China