Pal Hermansen / Brown bear (Ursus arctos), two showing aggression at night. Finland. August.
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Ingo Arndt / Grizzly bear (Ursus arctos horribilis) portrait, Lake Clark National Park, Alaska, USA, September.
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Carol Walker / Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA.
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Bence Mate / Great egrets (Ardea alba) fighting with necks intertwined, Lake Csaj, Kiskunsagi National Park, Hungary. January.
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Bence Mate / DUPLICATE Cuckoos (Cuculus canorus) squabbling, Kiskunsagi National Park, Pusztaszer, Hungary. February.
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Bence Mate / Montezuma oropendola (Psarocolius montezuma) two interacting, Costa Rica.
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Bence Mate / Great egrets (Ardea alba) sky pointing, sizing each other up before fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January.
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Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting, Kuusamo, Finland, April.
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Mark Bowler / Black capuchin, (Sapajus nigritus) female standing on hind legs, Rio de Janeiro, Brazil.
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Erlend Haarberg / Arctic fox (Alopex lagopus) cubs looking curiously, Dovrefjell-Sunndalsfjella National Park, Norway, July.
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Andy Sands / German wasp (Vespula germanica) worker carrying rubbish, flying out of nest in air brick, Hertfordshire, England, UK. July
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Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month cub, Ranthambhore National Park, India.