Michel Roggo / Chub fish (Leuciscus cephalus) and Mute swan (Cygnus color) swimming over, Rhine river, Schaffhausen, Switzerland. RUNNER-UP - Wildlife Photographer of the Year 2010.

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair on lake at sunset, North America

Flip de Nooyer / Mute Swan (Cygnus olor) three cygnets on parent's back, Europe

Flip de Nooyer / Mute Swan (Cygnus olor) pair courting, Europe

Konrad Wothe / Whooper Swan (Cygnus cygnus) pair arguing at wintering site, Kussharo-ko, Hokkaido, Japan

Konrad Wothe / Mute Swan (Cygnus olor) with three chicks riding on her back, Germany

Konrad Wothe / Mute Swan (Cygnus olor) courting pair, Germany

Jim Brandenburg / Tundra Swan (Cygnus columbianus) trio flying against cloudy sky, Minnesota

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair on lake, Copper River Delta, Alaska

Jim Brandenburg / Tundra Swan (Cygnus columbianus) flock taking off from lake, Mattamuskeet National Wildlife Reserve, North Carolina

Tui De Roy / Black-necked Swan (Cygnus melancoryphus) group swimming, Puerto Natales, Chilean Fjords, Chile

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair on lake, North America

Michael Quinton / Trumpeter Swan (Cygnus buccinator) parent and squabbling day-old cygnets, North America

Hiroya Minakuchi / Whooper Swan (Cygnus cygnus) group flying over Northern Pintail (Anas acuta) flock, Lake Hyo, Japan

Hiroya Minakuchi / Whooper Swan (Cygnus cygnus) and Northern Pintail (Anas acuta) flock, Lake Hyo, Japan

Donald M. Jones / Tundra Swan (Cygnus columbianus) group flying, Fairfield, Montana

Martin Willis / Black Swan (Cygnus atratus) swimming, Mongers Lake, Western Australia, Australia

Steve Gettle / Mute Swan (Cygnus olor) with cygnets, Kensington Metropark, Milford, Michigan

Steve Gettle / Mute Swan (Cygnus olor) with cygnets, Kensington Metropark, Milford, Michigan

Steve Gettle / Trumpeter Swan (Cygnus buccinator) swimming, Seney National Wildlife Refuge, Michigan

Steve Gettle / Mute Swan (Cygnus olor) flying, Kensington Metropark, Milford, Michigan

Steve Gettle / Trumpeter Swan (Cygnus buccinator) on nest, Wyoming

Yva Momatiuk & John Eastcott / Black Swan (Cygnus atratus) flock, Lake Rotorua, New Zealand

Yva Momatiuk & John Eastcott / Black Swan (Cygnus atratus), Lake Rotorua, New Zealand

Yva Momatiuk & John Eastcott / Black Swan (Cygnus atratus) pair, Lake Rotorua, New Zealand

Konrad Wothe / Mute Swan (Cygnus olor) pair courting, Bavaria, Germany

Donald M. Jones / Tundra Swan (Cygnus columbianus) flying, central Montana

Donald M. Jones / Tundra Swan (Cygnus columbianus) flock flying, central Montana

Pete Oxford / Black-necked Swan (Cygnus melancoryphus), Keppel Island, Falkland Islands

Pete Oxford / Black-necked Swan (Cygnus melancoryphus), Keppel Island, Falkland Islands

Pete Oxford / Black-necked Swan (Cygnus melancoryphus), Keppel Island, Falkland Islands

Michael Quinton / Trumpeter Swan (Cygnus buccinator) standing in pond, Alaska

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair swimming, Alaska

Jasper Doest / Mute Swan (Cygnus olor), South Holland, Netherlands

Jasper Doest / Whooper Swan (Cygnus cygnus) pair flying at sunset, Sweden

Stephen Belcher / Whooper Swan (Cygnus cygnus) flock swimming on lake with steam, Lake Kussharo-ko, Hokkaido, Japan

Stephen Belcher / Whooper Swan (Cygnus cygnus) flock swimming on lake with steam, Lake Kussharo-ko, Hokkaido, Japan

Stephen Belcher / Whooper Swan (Cygnus cygnus) group flying, Lake Kussharo-ko, Hokkaido, Japan

Stephen Belcher / Whooper Swan (Cygnus cygnus) pair standing on frozen lake, calling, Lake Kussharo-ko, Hokkaido, Japan

Sebastian Kennerknecht / Tundra Swan (Cygnus columbianus) pairs fighting, Tule Lake National Wildlife Refuge, California

Sebastian Kennerknecht / Tundra Swan (Cygnus columbianus) landing, Tule Lake National Wildlife Refuge, California

Sebastian Kennerknecht / Tundra Swan (Cygnus columbianus) pair landing, Tule Lake National Wildlife Refuge, California

Colin Monteath / Black Swan (Cygnus atratus), Canterbury, New Zealand

Colin Monteath / Black Swan (Cygnus atratus), Canterbury, New Zealand

Jasper Doest / Whooper Swan (Cygnus cygnus) flapping its wings, Sweden

Jan Vermeer / Whooper Swan (Cygnus cygnus) landing on the ice, Hokkaido, Japan

Mitsuaki Iwago / Australian Spiny Lobster (Panulirus cygnus) on a moss-covered rock, Australia

Jan Vermeer / Whooper Swan (Cygnus cygnus) courting in frozen wintering areas of Hokkaido, Japan

Michio Hoshino / Tundra Swan (Cygnus columbianus) flock flying, Alaska

Konrad Wothe / Mute Swan (Cygnus olor) flying, Camargue, France

Konrad Wothe / Mute Swan (Cygnus olor) on lake, Bavaria, Germany

Yva Momatiuk & John Eastcott / Black-necked Swan (Cygnus melancoryphus) near Puerto Natales, Patagonia, Argentina

Yva Momatiuk & John Eastcott / Black-necked Swan (Cygnus melancoryphus) near Puerto Natales, Patagonia, Argentina

Pete Oxford / Mute Swan (Cygnus olor) attacking Mallard (Anas platyrhynchos) ducklings, Wyoming

Thomas Marent / Mute Swan (Cygnus olor) pair courting, Switzerland

Thomas Marent / Mute Swan (Cygnus olor) male threat displaying, Switzerland

Thomas Marent / Black Swan (Cygnus atratus), Switzerland

Thomas Marent / Whooper Swan (Cygnus cygnus) portrait, Switzerland

Thomas Marent / Mute Swan (Cygnus olor) coming in for a landing, Switzerland

Thomas Mangelsen / Trumpeter Swan (Cygnus buccinator) group walking along the unfrozen Chilkat River, Alaska

Ingo Arndt / Whooper Swan (Cygnus cygnus) group feeding in marsh, Norway

Konrad Wothe / Whooper Swan (Cygnus cygnus) pair flying, Germany

Konrad Wothe / Mute Swan (Cygnus olor) adult incubating eggs on nest on lake, Upper Bavaria, Germany

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair on lake at sunset, North America

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair on lake at sunset, North America

Jim Brandenburg / Trumpeter Swan (Cygnus buccinator) silhouetted in mist, Minnesota

Jim Brandenburg / Trumpeter Swan (Cygnus buccinator) silhouetted in mist, Minnesota

Flip de Nooyer / Black Swan (Cygnus atratus) parent swimming with seven white cygnets, Europe

Flip de Nooyer / Mute Swan (Cygnus olor) courting pair on lake, Europe

Winfried Wisniewski / Whooper Swan (Cygnus cygnus) tagged for research, Japan

Winfried Wisniewski / Whooper Swan (Cygnus cygnus) stretching at sunset, Japan

Winfried Wisniewski / Whooper Swan (Cygnus cygnus) foraging in lake, Japan

Winfried Wisniewski / Whooper Swan (Cygnus cygnus) landing in water, Japan

Winfried Wisniewski / Whooper Swan (Cygnus cygnus) close up of head, Japan

Flip de Nooyer / Mute Swan (Cygnus olor) parent on nest with cygnets, Europe

Duncan Usher / Mute Swan (Cygnus olor) pair flying, Europe

Winfried Wisniewski / Whooper Swan (Cygnus cygnus) pair interacting, ranges in the palearctic, winters to Indian and southeastern China

Winfried Wisniewski / Whooper Swan (Cygnus cygnus) flying, ranges in the palearctic, winters to Indian and southeastern China

Jan Vermeer / Mute Swan (Cygnus olor) flock in water

ZSSD / Whooper Swan (Cygnus cygnus) portrait

Flip de Nooyer / Black Swan (Cygnus atratus) cygnet, portrait, front view

Flip de Nooyer / Mute Swan (Cygnus olor) three cygnets on the back of parent, Europe

Flip de Nooyer / Mute Swan (Cygnus olor) pair courting, Europe

Flip de Nooyer / Mute Swan (Cygnus olor) pair courting, Europe

Matthias Breiter / Tundra Swan (Cygnus columbianus) on frozen pond, Alaska Range, Tok, Alaska

Flip de Nooyer / Mute Swan (Cygnus olor) two adults, Europe

Flip de Nooyer / Mute Swan (Cygnus olor) cygnet under its parent's wing, Europe

Flip de Nooyer / Mute Swan (Cygnus olor) cygnet under parent's wing, Europe

Flip de Nooyer / Mute Swan (Cygnus olor) large flock, Europe

Yva Momatiuk & John Eastcott / Tundra Swan (Cygnus columbianus) large flock flying over Mattamuskeet National Wildlife Refuge, winter, North Carolina

Konrad Wothe / Whooper Swan (Cygnus cygnus) group on icy lake at wintering site, Kussharo-ko, Hokkaido, Japan

Konrad Wothe / Mute Swan (Cygnus olor) flying, Germany

Konrad Wothe / Whooper Swan (Cygnus cygnus) pair wintering, Hokkaido, Japan

Thomas Mangelsen / Trumpeter Swan (Cygnus buccinator) flock flying at sunset, North America

Thomas Mangelsen / Trumpeter Swan (Cygnus buccinator) flock flying in formation, Snake River, Nebraska

Konrad Wothe / Whooper Swan (Cygnus cygnus) and Northern Pintail (Anas acuta) mixed flock, Odaito, Hokkaido, Japan