Jack Dykinga / Lightning storm with rainbow over Saguaro cactus (Carnegiea gigantea) near Redrock, Arizona State Trust, Sonoran Desert, Arizona. September 2015. Long exposure with lightning trigger.

Martin Dohrn / Herd of giraffes (Giraffa camelopardalis) resting at night, Masai Mara, Kenya. Image taken at night using thermal camera technology, with no artificial light.

Eduardo Blanco / Pale burrower mayfly (Ephoron virgo), swarming in the millions on street at night, appearing like snow. Mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August 2019. Finalist i...

Doug Gimesy / Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016. COP26 Countdown Photo Competi...

Doug Gimesy / Little penguin (Eudyptula minor) on rocks of St Kilda breakwater, Melbourne city lights in background. Victoria, Australia. January 2017.

David Tipling / Barn owl(Tyto alba) flying, North Norfolk, England, UK. february

Guy Edwardes / Aurora Borealis over Sycamore Gap, Hadrian's Wall, Northumberland, England. February 2014

Guy Edwardes / Northern Lights over Vestrahorn, Hofn, Iceland. September 2017

Solvin Zankl / Bioluminescent Sea-fireflies (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen and is for mating displays or defenc...

Ingo Arndt / View from Margaret Knoll lookout of sunset and rain clouds over the Indian Ocean and the east coast of Christmas Island, Indian Ocean, Australian Territory, December 2009.

Wim van den Heever / Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April.

Claudio Contreras / Light entering Cenote Siete Bocas, near Puerto Morelos, Riviera Maya, Yucatan Peninsula, Mexico. January

Bence Mate / Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night. Winner of the Mammals Category of the GDT Awards 20...

Pete Oxford / Woman inside Nyshi Long House, Nyshi Tribe, Arunachal Pradesh.North East India, November 2014.

Neil Aldridge / Urban Red fox (Vulpes vulpes) searching for food near bins in a courtyard at night, West London, United Kingdom, May 2014. Finalist in the Urban category of the Wildlife Photographer of the Year Awar...

Sven Zacek / Eilean Donan Castle lit up at twilight with dark clouds. Scotland, UK, September 2013.

Juan Carlos Munoz / Tourists looking down onto underground stream, from illuminated pathway, Skocjan Caves, Green Karst, Slovenia, October 2014.

Yukihiro Fukuda / Light trails of Japanese fireflies (Hotaria parvula and Luciola cruciata), Nichinan-chou, Tottori, Japan, July, Endemic species.

Juergen Freund / Wangi Falls at night illuminated by moonlight, Litchfield National Park, Northern Territory, Australia. December 2012.

Roy Mangersnes / Full moon over Adventsdlen, Longyearbyen, Svalbard, Norway, April.

Denis-Huot / African elephant (Loxodonta africana) herd at sunset, Masai-Mara Game Reserve, Kenya. January.

David Noton / Aurora Borealis (Northern Lights) over the Klondike River, Yukon Territories, Canada, September 2013.

Sam Hobson / Young urban Red fox (Vulpes vulpes). Bristol, UK. August.

Olga Kamenskaya / Ice stalagmites / ice spikes formation, Lake Baikal, Siberia, Russia, March.

Jouan Rius / Antelope Canyon, a slot canyon with eroded sandstone patterns, Navajo Tribal Park, Arizona, USA, December 2012.

Jouan Rius / Gypsum sand dune at sunset, White Sands National Park, Chihuahuan Desert, New Mexico, USA, December 2012.

Nature Production / Japanese firefly (Luciola cruciata) on leaf at night, with light trails of hundreds of other fireflies in flight in the background, Japan

Alex Mustard / Scenic view of a bull kelp forest (Nereocystis luetkeana) with sunlight shining through the fronds. Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean.

Andy Rouse / Maasai Giraffe (Giraffa camelopardalis tippelskirchi) under setting sun Maasai Mara, Kenya, Africa

Jouan Rius / Sandstone cliff at sunset, Colorado Plateau, Kodachrome Basin State Park, Utah, USA November 2012

Theo Bosboom,Theo Bosboom / Abstract reflections of reed and clouds in the water, Texel, the Netherlands, April

Juan Carlos Munoz / People descending into the Thrihnukagigur volcano a dormant volcano, now used for tours, Iceland, Europe, June 2012

Sandra Bartocha / Cowslip (Primula veris) and Cuckoo Flower (Cardamine pratensis) in flower with light effects, Brandenburger Vorstadt, Potsdam, Germany, April

Ben Hall / 2020VISION / Sun's rays shining through dark clouds, Ballynahone Bog, County Antrim, Northern Ireland, UK, June 2011

Fabrice Cahez / Sunrise breaking though dense woodland canopy. Vosges forest, France, September 2010.

Martin Dohrn / Three hyenas (Crocuta crocuta) approaching a resting lion (Panthera leo) at night. Topi Plain, Masai Mara, Kenya. Image taken using an infared camera with no artificial visible light, on location for...

Martin Dohrn / Group of hyena (Crocuta crocuta) playing in deep rain puddle at night. Topi Plain, Masai Mara, Kenya. Image taken using "starlight camera" with no artificial light, on location for National Geographi...

Martin Dohrn / Thermal camera image of hippos (Hippopotamus amphibius), crocodile (Crocodylus niloticus) and wildebeest (Connochaetes taurinus, far left) on the banks of the Mara River, Masai Mara, Kenya. Image take...

Inaki Relanzon / Dramatic light on Benijo Beach. Anaga Peninsula, Tenerife, Canary Islands, February 2011.

Andy Rouse / Grizzly Bear (Ursus arctos) on the move in late evening light after heavy rain. Alaska, USA, June.

Martin Dohrn / Sleeping African elephants (Loxodonta africana), two adults and offspring, Masai Mara, Kenya. Image taken at night using thermal camera technology without artificial light.

Floris van Breugel / Joshua Trees (Yucca brevifolia) in Antelope Valley against the night sky illuminated by the orange glow from Los Angeles light pollution, California, USA. September 2009.

Edwin Giesbers / Snow Leopard (Panthera uncia) sitting in darkness, with eyes glowing as they reflect available light, captive

Mark Bowler / Fireflies {Lampyridae} in rainforest, Singapore.

Martin Dohrn / The Southern Cross star constellation shining above Ballenites trees in Masai Mara, Kenya. Starlight image intensifier camera image taken with no artificial light.

Martin Dohrn / Spotted hyaenas at night, Masai Mara, Kenya. Starlight image intensifier camera image taken with no artificial light, composite

Martin Dohrn / Bioluminescence - pool with Snook, Snapper and a Stingray fish, Bio Bay, Vieques Island, Puerto Rico. Central America. Starlight image intensifier camera image taken with no artificial light.

George Sanker / Meander of Gardiner River with grassland and hills in background, in evening light. Yellowstone National Park, Wyoming, USA. June 2018.

Franco Banfi / Camp on beach at full moon, in Vrangel Bay where Bowhead whale (Balaena mysticetus) congregate during summer. Primorsky Krai, Russia. August 2019.

Eduardo Blanco / Pale burrower mayfly (Ephoron virgo) swarming around streetlights at night, with gecko (Tarentola mauritanica) inside lamp. Mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. Aug...

Doug Gimesy / Researcher from platypusSPOT checking fyke net for Platypus (Ornithorhynchus anatinus), part of a river population study for Melbourne Water. Beaconsfield, Melbourne, Victoria, Australia. 2017.

Guy Edwardes / Beech (Fagus sylvatica) woodland in moonlight. Cranborne Chase, Dorset, England, UK. March 2019.

Yashpal Rathore / Leopard cat (Prionailurus bengalensis) walking along ridge at night. Lights on hills in background. Nilgiri Biosphere Reserve, India. 2019. Camera trap image.

Yashpal Rathore / Brown palm civet (Paradoxurus jerdoni) on hillside, lights of rural settlements in background. Nilgiri Biosphere Reserve, India. Camera trap image.

Juan Carlos Munoz / Coastal town of Saint-Jean-de-Luz, view across sea. Pyrenees Atlantiques, Nouvelle-Aquitaine, France. December 2014.

Ingo Arndt / Sunbeams over Lago Sarmiento. Torres del Paine, Patagonia, Chile. December 2016.

Doug Gimesy / Earthcare St Kilda penguin research volunteer holding Little penguin (Eudyptula minor) after retrieving it from burrow to check for microchip and determine sex and weight. St Kilda breakwater, Melbour...

Doug Gimesy / Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016.

Doug Gimesy / Researchers from Earthcare St Kilda preparing to microchip a moulting Little penguin (Eudyptula minor). Bill diameter will be measured to determine sex. St Kilda breakwater, Melbourne, Victoria, Austr...

Doug Gimesy / Researchers from Earthcare St Kilda scanning a moulting Little penguin (Eudyptula minor) for microchip. St Kilda breakwater, Melbourne, Victoria, Australia. December 2016. Model released.

Doug Gimesy / Earthcare St Kilda penguin research volunteers with Little penguin (Eudyptula minor) checking for microchip and determining sex and weight. St Kilda breakwater, Melbourne, Victoria, Australia. January...

Doug Gimesy / Little blue penguin (Eudyptula minor) on St Kilda breakwater, lights of Melbourne in background. Victoria, Australia. November 2016.

Doug Gimesy / Little blue penguins (Eudyptula minor), two standing on rocks of St Kilda breakwater. Melbourne city lights in the background. Victoria, Australia. December 2016.

Doug Gimesy / Earthcare St Kilda penguin research volunteers with Little penguin (Eudyptula minor) checking for microchip and determining sex and weight. St Kilda breakwater, Melbourne, Victoria, Australia. January...

Doug Gimesy / Earthcare St Kilda penguin research volunteers with Little penguin (Eudyptula minor) checking for microchip and determining sex and weight. St Kilda breakwater, Melbourne, Victoria, Australia. Novembe...

Enrique Lopez-Tapia / Man with lantern lighting up skulls on rock at Toraja cemetery. The Toraja culture of West and South Sulawesi revolves around death with funeral ceremonies an important part of daily life. Indonesia....

Enrique Lopez-Tapia / Man looking at skull, coins and cigarettes in cave, Toraja cemetery. The Toraja culture of West and South Sulawesi revolves around death with funeral ceremonies an important part of daily life. Indone...

Heather Angel / Love-in-a-mist (Nigella damascena) stamens fluroescing in UV light. Ring of nectaries are dark red with blue UV reflective hairs.

Nick Upton / Radio-collared Male Pine Marten (Martes martes) emerging from a temporary soft release cage after dark during the Forest of Dean and River Wye Pine Marten Project, the Forest of Dean, Gloucestershire,...

Nick Upton / Radio-collared Male Pine Marten (Martes martes) emerging from a temporary soft release cage after dark during the Forest of Dean and River Wye Pine Marten Project, the Forest of Dean, Gloucestershire,...

Nick Upton / Radio-collared Male Pine Marten (Martes martes) emerging from a temporary soft release cage after dark during the Forest of Dean and River Wye Pine Marten Project, the Forest of Dean, Gloucestershire,...

David Tipling / European hedgehog (Erinaceus europaeus) using hole in garden fence to move between gardens, Norfolk, England, UK.

SCOTLAND: The Big Picture / Light trails from car lights, evening rush hour in Glasgow city centre. Scotland, UK.

SCOTLAND: The Big Picture / Evening rush hour with trails from car lights, long exposure. Charing Cross, Glasgow, Scotland, UK.

Guy Edwardes / Aurora Borealis over Sycamore Gap, Hadrian's Wall, Northumberland, England. February 2014

Guy Edwardes / Northern Lights over Reyjanes Pensinsula, Iceland. September 2017

Ashley Cooper / Young boy doing homework in house lit by electricity from battery. Batteries are solar charged from WWF project to supply electricity to a remote island in the Sunderbans, Ganges Delta, India. Decembe...

Ashley Cooper / The Lloyds insurance building in the City of London, England, UK, December 2009.

Ashley Cooper / Office blocks lit up at night with Hong Kong, China. February 2010.

Ashley Cooper / Office blocks lit up at night and cars in Hong Kong, China. February 2010.

Ashley Cooper / Office blocks lit up at night in Hong Kong, China. February 2010.

Ashley Cooper / Office blocks lit up at night with Hong Kong, China. February 2010.

Ashley Cooper / Office blocks lit up at night with Hong Kong, China. February 2010.

Solvin Zankl / Bioluminescent Firefly squid (Watasenia scintillans) emits intense flashes of light from three tiny luminous organs (photophores) that are located at the tip of each of a pair of ventral arms, as wel...

Solvin Zankl / Bioluminescent Firefly squid (Watasenia scintillans) emitting intense flashes of light from three tiny luminous organs (photophores) that are located at the tip of each of a pair of ventral arms, as...

Solvin Zankl / Bioluminescent Sea-fireflies (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen and is for mating displays or defenc...

Solvin Zankl / Bioluminescent Sea-fireflies (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen and is for mating displays or defenc...

Solvin Zankl / Bioluminescent Sea-fireflies (Vargula hilgendorfii) washed up on shore, producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen and is for mati...

Solvin Zankl / Bioluminescent Sea-firefliy (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen and is for mating displays or prey de...

Solvin Zankl / Firefly squid (Watasenia scintillans) emitting light from photophores, Toyama Bay, Japan. This form of bioluminescence is used as counter illumination camouflage. See image 1601190 for comparison with...

Doug Perrine / Comb jelly / sea gooseberry (Hormiphora palmata) diffracting light through cilia (fine hair-like structures) on the comb rows, Kailua Kona, Hawaii, USA.

Doug Perrine / Close-up of Ctenes or Comb plates (Eurhamphaea vexilligera) a translucent gelatinous organism with colors produced by the diffraction of light through cilia. Kailua Kona, Hawaii, USA.

Bruno D'Amicis / Light filtering through clouds on rugged southern slope of Mount Prena in the Gran Sasso massif. Gran Sasso NP, Central Apennines, Abruzzo, Italy, September.

Michel Roggo / Under water in the Abismo Anhumas or Anhumas Abyss. This is a 80 metre deep lake, at the bottom of a 72 metre deep cave. Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul...

Michel Roggo / Diver underwater in the Abismo Anhumas or Anhumas Abyss. This is a 80 metre deep lake, at the bottom of a 72 metre deep cave. Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso d...

Michel Roggo / Under water in the Abismo Anhumas or Anhumas Abyss. This is a 80 metre deep lake, at the bottom of a 72 metre deep cave. Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul...