Bence Mate / Great egrets (Ardea alba) and Grey heron (Ardea cinerea) squabbling over fish, with some in flight. Blurred motion photograph, Lake Csaj, Pusztaszer, Hungary. February.

Edwin Giesbers / Scarlet macaw (Ara macao) flying, blurred motion. Costa Rica.

Chris & Monique Fallows / Indo Pacific Sailfish (Istiophorus platypterus) feeding on sardines (Sardina pilchardus), Isla Mujeres, Mexico, March.

Dr. Axel Gebauer / Oxeye daisy (Leucanthemum vulgare) abstract, Upper Lusatia Heath and Pond Landscape, Upper Lusatia UNESCO Biosphere Reserve, Germany

Denis-Huot / Spotted hyena (Crocuta crocuta) hunting Lesser flamingoes (Phoenicopterus minor) Lake Nakuru, Kenya

Andrew Walmsley / Arctic terns (Sterna paradisaea), adults defending their nests from photographers, Northumberland, June. No release available.

Ernie Janes / Greylag geese (Anser anser) in flight with paraglider behind, Cley bank, Norfolk, UK

Alex Mustard / A bull California sealion (Zalophus californianus) bursting through a school of baitfish in the shallows. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean.

Roy Mangersnes / Aerial view of Gentoo Penguin (Pygoscelis papua) rookery on Cuverville Island, Antarctica. November. The birds are in panic as a helicopter passes by.

Nick Upton / Dr. Jenny Macpherson carryng a live trap containing a Pine Marten (Martes martes) caught at night in Scottish woodland for a reintroduction project to Wales run by the Vincent Wildlife Trust, Highland...

Nick Upton / Dr. Jenny Macpherson entering a soft release cage at night carrying a Pine Marten (Martes martes) in a transport crate on arrival from Scotland, during a reintroduction project by the Vincent Wildlife...

Nick Upton / Dr. Jenny Macpherson inspecting Pine Martens (Martes martes) in an animal transport van at a motorway servce station en route from Scotland to Wales for a reintroduction project run by the Vincent Wil...