Francisco Marquez / Round up of wild mares and foals from marshland to the corrals of the town. Donana National Park, Spain. September 2014.

Ramon Navarro / Whiskered tern (Chlydonias hybrida) passing frog prey to a chick at nest, Donana National Park, Seville, Spain, August.

Kristel Richard / A herd of Retuerta mares trotting, Guadiamar scientific reserve, Donana National Park, Andalusia, Spain. This breed of horse was close to extinction in the 1990s with only 5 individuals left, but now...

Roland Seitre / Iberian / Spanish Lynx (Lynx pardina) cub, 2-3 months, in transport box at the Donana breeding station. Lynx reintroduction program, Andalusia, Spain, June 2007.

Roland Seitre / Program director Dr Astrid Vargas examining Iberian / Spanish Lynx (Lynx pardina) at the Donana breeding station. Andalusia, Spain, June 2007.

Roland Seitre / Mother Iberian / Spanish Lynx (Lynx pardina) with cubs climbing the fence of their enclosure. Donana breeding station, part of the lynx reintroduction project in Andalusia. Spain, June 2006.

Juan Carlos Munoz / Aquatic plants in water, Donana NP, Andalucia, Spain

Angelo Gandolfi / Short-toed Eagle (circaetus gallicus) flying amongst Stone Pines (Pinus pinea). Coto Donyana, Andalusia, Spain. 2007.

Jose B. Ruiz / White storks (Ciconia ciconia) roosting in tree at night, Donana NP, Spain

Jose B. Ruiz / Field of cultivated Rice (Oryza sativa) at dawn, Donana NP, Spain

Jose B. Ruiz / Silhouette of White storks (Ciconia ciconia) in tree at dawn, Donana NP, Spain

Jose B. Ruiz / Silhouette of White stork (Ciconia ciconia) flying from tree at dawn, Donana NP, Spain

Jose B. Ruiz / Female Black winged stilt {Himantopus himantopus} with summer plumage, foraging in lake, Donana NP, Sevilla, Spain

Jose B. Ruiz / Grey heron {Ardea cinerea} wading in lake, Donana NP, Sevilla, spain

Jose B. Ruiz / Black winged stilt {Himantopus himantopus} with head submerged, feeding in lake, Donana NP, Sevilla, Spain

Jose B. Ruiz / Little Egret {Egretta garzetta} portrait wading and looking for prey in water, Donana NP, Sevilla, Spain

Jose B. Ruiz / Little Egret {Egretta garzetta} making a splash whilst hunting for food in lake, Donana NP, Sevilla, Spain

Jose B. Ruiz / Little egret {Egretta garzetta} at waters edge hunting sequence 1/3, Donana NP, Sevilla, Spain

Jose B. Ruiz / Little egret {Egretta garzetta} at waters edge hunting sequence 2/3, Donana NP, Sevilla, Spain

Jose B. Ruiz / Little Egret {Egretta garzetta} making a splash whilst hunting for food in lake, Donana NP, Sevilla, Spain.

Jose B. Ruiz / Whiskered tern {Chlydonias hybrida} in flight with food for chicks, Donana NP, Sevilla, Spain

Jose B. Ruiz / Glossy ibis {Plegadis falcinellus} dispalying in wetland, sequence 2/2, Donana NP, Spain

Jose B. Ruiz / Little Egret {Egretta garzetta} portrait, Donana NP, Sevilla, Spain

Jose B. Ruiz / Black tailed godwits {Limosa limosa} landing on water surface, Donana NP, Sevilla, Spain

Jose B. Ruiz / Black winged stilt {Himantopus himantopus} Donana NP, Sevilla, Spain

Jose B. Ruiz / Spanish / Iberian Lynx in Donana NP, Spain

Jose B. Ruiz / Spanish /Iberian Lynx in Donana NP, Spain

Jose B. Ruiz / Spanish / Iberian Lynx in Donana NP, Spain (Lynx pardina)

Ashley Cooper / Snails aestivating on a fence post in the Coto Donana, Andalucia, Spain, June 2011

Ashley Cooper / White Storks (Ciconia ciconia) nesting on electricity pylons in the Coto Donana in Andalucia, Spain. June 2011

Francisco Marquez / Horse riders relaxing after rounding up mares and foals. Donana National Park, Spain. September 2014.

Francisco Marquez / Herd of horses driven to the hermitage of El Rocio to be blessed. Coto Donana, Spain. June 2014.

Francisco Marquez / Herd of horses driven to the hermitage of El Rocio to be blessed. Coto Donana, Spain. June 2014.

Francisco Marquez / Horses drinking from trough after round up, Donana National Park, Spain. September 20014.

Francisco Marquez / Carts carrying food, drink and beds to the fields where the horsemen work, rounding up mares and foals. Donana National Park, Spain. June 2014.

Francisco Marquez / Round up of mares and foals, Donana National Park, Spain. September 2014.

Francisco Marquez / Man on horseback during the round up and transfer of the mares and foals from the marsh to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Men on horseback during the annual round up and transfer of the mares and foals from the marsh to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Man on horseback during the annual round up and transfer of the mares and foals from the marsh to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Men on horseback during the round up of mares and foals on marshland, with rainbow. Donana National Park, Spain. September 2014.

Francisco Marquez / Man on horseback during the annual capture and transfer of the mares and foals from the marsh to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Round up of mares and foals from marshland to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Men on horseback during the round up of mares and foals from marshland to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Man on horseback during the round up of mares and foals from marshland to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Round up of wild mares and foals from marshland to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Man on horseback during the round up of mares and foals from marshland to the corrals of the town. Donana National Park, Spain. September 2014.

Francisco Marquez / Close up of branded mark on horse. Mares and foals are caught into corrals where each yeguero (owner of the mares) identifies their animals, separates the foals from their mothers, worm them, cut the...

Francisco Marquez / Branding irons heating in fire. Mares and foals are caught into corrals where each yeguero (owner of the mares) identifies their animals, separates the foals from their mothers, worm them, cut their...

Francisco Marquez / Man cutting foal's mane. During the round up of mares and foals are caught into corrals where each yeguero (owner of the mares) identifies their animals, separates the foals from their mothers, worm t...

Francisco Marquez / Man cutting foal's mane. During the round up of mares and foals are caught into corrals where each yeguero (owner of the mares) identifies their animals, separates the foals from their mothers, worm t...

Francisco Marquez / Man with scissors for cutting foal's mane. During the round up of mares and foal's are caught into corrals where each yeguero (owner of the mares) identifies their animals, separates the foals from t...

Francisco Marquez / Mares and foals in corral, with man pulling mare towards him. During the round up of mares and foals each yeguero (owner of the mares) identifies their animals, separates the foals from their mothers,...

Francisco Marquez / Foals are driven into corrals where each yeguero (owner of the mares) i identifies their animals, separates the foals from their mothers, worm them, cut their mane and brand them before sale. Donana N...

Francisco Marquez / Foals are driven into corrals where each yeguero (owner of the mares) identifies their animals, separates the foals from their mothers, worm them, cut their mane and brand them before sale. Donana Nat...

Francisco Marquez / Close up of horses nose peering out of corral. During the round up of mares and foals each yeguero (owner of the mares) identifies their animals, separates the foals from their mothers, worm them, cut...

Francisco Marquez / Mares and foals rounded up into corral. During the annual round up of mares and foals. Donana National Park, Spain.. September 2014.

Francisco Marquez / Horse riders relaxing after rounding up mares and foals. Donana National Park, Spain. September 2014.

Francisco Marquez / Horse riders relaxing after rounding up mares and foals. Donana National Park, Spain. September 2014.

Andres M. Dominguez / Short-eared owl (Asio flammeus) perched on a fence post, Doñana Natural Park, Andalusia, Spain. January.

Andres M. Dominguez / Short-eared owl (Asio flammeus) perched, Donana Natural Park, Andalusia, Spain. January.

Andres M. Dominguez / Short-eared owl (Asio flammeus) perched and preening feathers, Donana Natural Park, Andalusia, Spain. January.

Andres M. Dominguez / Short-eared owl (Asio flammeus) perched and preening feathers, Donana Natural Park, Andalusia, Spain. January.

Andres M. Dominguez / Short-eared owl (Asio flammeus) camouflaged, Donana Natural Park, Andalusia, Spain. January.

Andres M. Dominguez / Short-eared owl (Asio flammeus) camouflaged, Donana Natural Park, Andalusia, Spain. January.

Andres M. Dominguez / Black shouldered kite (Elanus caeruleus) perched on branch, yawning, Donana Natural Park, southern Spain, December.

Ernie Janes / Bark of Stone pine (Pinus pinea) Coto Donana, Spain.

Ernie Janes / Stone pine (Pinus pinea) trees, Coto Donana, Spain.

Ernie Janes / Board walk in Donana National Park, Almonte, Spain, October 2017.

Andres M. Dominguez / Short-eared owl (Asio flammeus) perched on a fence post, Donana Natural Park, Andalusia, Spain. January.

Andres M. Dominguez / Female Kestrel (Falco tinnunculus) in flight, Donana Natural Park, Andalusia, Spain. January.

Francisco Marquez / Red deer (Cervus elaphus) fawn hidden in long grass, Donana National Park, Andalusia, Spain, August.

Juan Carlos Munoz / Common snipe (Gallinago gallinago) hidden in vegetation, Donana National Park, Huelva, Andalusia, Spain. February.

Andres M. Dominguez / Greater flamingo (Phoenicopterus roseus) with El Rocio Church silhouetted at sunset, Donana National Park, southern Spain, November 2009.

Pascal Tordeux / Horse walking through wetlands, Donana, Andalusia, Spain, March 2014.

Pascal Tordeux / Red deer (Cervus elaphus) stags in Donana National Park, Andalusia, Spain, March.

Pascal Tordeux / Traditional thatched hut, Donana National Park, Andalusia, Spain, March 2014.

Pascal Tordeux / Park ranger on horseback next to the Guadalquivir river, Donana National Park, Andalusia, Spain, March 2014.

Pascal Tordeux / Greater flamingos (Phoenicopterus roseus) in wetland habitat, Donana National Park, Andalusia, Spain, March.

Pascal Tordeux / Pine trees in the fog near Las Marismillas Palace, Donana National Park, Andalusia, Spain, March 2014.

Pascal Tordeux / Pine tree (Pinus sp) on the edge of sand dune, Guadalquivir valley, Donana National Park, Andalusia, Spain, March 2014.

Pascal Tordeux / Sand dunes, Donana National Park, Andalusia, Spain, March 2014.

Pascal Tordeux / Pine trees (Pinus sp) partially buried by sand dune, Donana National Park, Andalusia, Spain, March 2014.

Pascal Tordeux / Rippled sand dune, Donana National Park, Andalusia, Spain, March 2014.

Pascal Tordeux / Sand dunes, Donana National Park, Andalusia, Spain, March 2014.

Pascal Tordeux / Greater flamingos (Phoenicopterus roseus) in flight over sand dune, Donana National Park, Andalusia, Spain, March.

Pascal Tordeux / Pine trees (Pinus sp) at risk of being buried by the shifting sand dunes, Donana National Park, Andalusia, Spain, March 2014.

Ramon Navarro / Female Common kingfisher (Alcedo atthis) perched on a branch, preening, Donana National Park, Seville, Spain, October.

Ramon Navarro / Whiskered tern (Chlydonias hybrida) passing insect prey to a chick at nest, Donana National Park, Seville, Spain, August.

Ramon Navarro / Whiskered tern (Chlydonias hybrida) passing insect prey to a chick at nest, Donana National Park, Seville, Spain, August.

Ramon Navarro / Whiskered tern (Chlidonias hybrida) perched on a tree stump, preening, Donana National Park, Seville, Spain, August.

Ramon Navarro / Whiskered tern (Chlydonias hybrida) settling on nest to incubate chicks, Donana National Park, Seville, Spain, August.

Ramon Navarro / Whiskered tern (Chlydonias hybrida) passing frog prey to a chick at nest, Donana National Park, Seville, Spain, August.

Ramon Navarro / Whiskered tern (Chlydonias hybrida) settling on nest to incubate chicks, Donana National Park, Seville, Spain, August.

Ramon Navarro / Pair of Whiskered terns (Chlydonias hybrida) at nest with chicks, with one chick eating a frog, Donana National Park, Seville, Spain, August.

Ramon Navarro / Pair of Whiskered terns (Chlydonias hybrida) at nest with chicks, Donana National Park, Seville, Spain, August.

Juan Carlos Munoz / Mixed flock of White storks (Ciconia ciconia), Common spoonbills (Platalea leucorodia) and Cattle egrets (Bubulcus ibis) settling to roost in a tree, Doñana National Park, Andalusia, Spain, May.