Michio Hoshino / Galapagos Giant Tortoise (Chelonoidis nigra) pair communicating, Galapagos Islands, Ecuador

Tui De Roy / Galapagos Giant Tortoise (Chelonoidis nigra) yawning, Alcedo Volcano, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Saddleback Galapagos Tortoise (Chelonoidis nigra hoodensis) old male portrait, Pinzon Island, Galapagos Islands, Ecuador

Pete Oxford / Saddleback Galapagos Tortoise (Chelonoidis nigra hoodensis) foot, Wolf Volcano, Isabella Island, Galapagos Islands, Ecuador

Pete Oxford / Galapagos Giant Tortoise (Chelonoidis nigra) foot, Wolf Volcano, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) wallowing in mud, Isabella Island, Galapagos Islands, Ecuador

Pete Oxford / Indefatigable Island Tortoise (Chelonoidis nigra porteri) grazing in the highlands, Santa Cruz Island, Galapagos Islands, Ecuador

Pete Oxford / Indefatigable Island Tortoise (Chelonoidis nigra porteri) highlands, Santa Cruz Island, Galapagos Islands, Ecuador

Ingo Arndt / Galapagos Giant Tortoise (Chelonoidis nigra) portrait, Isabella Island, Galapagos Islands, Ecuador

Ingo Arndt / Galapagos Giant Tortoise (Chelonoidis nigra) portrait, Isabella Island, Galapagos Islands, Ecuador

Ingo Arndt / Galapagos Giant Tortoise (Chelonoidis nigra) in grass, Santa Cruz Island, Galapagos Islands, Ecuador

Ingo Arndt / Galapagos Giant Tortoise (Chelonoidis nigra) close up of foot, Santa Cruz Island, Galapagos Islands, Ecuador