Benjamin Barthelemy,Benjamin Barthelemy / Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June
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David Noton / Palm trees line the beach at Pigeon Point, Tobago
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Tony Heald / Hippopotamus {Hippopotamus amphibius} territorial defense at water edge, Chobe national park, Botswana.
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Oscar Dewhurst / Red fox (Vulpes vulpes) sitting on a frozen lake at the edge of the reeds. London, UK. December.
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Markus Varesvuo / Hazel grouse (Tetrastes bonasia) standing in snow at forest edge. Helsinki, Finland. February.
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Axel Gomille / Wolf (Canis lupus), cubs playing at edge of woodland, Saxony-Anhalt, Germany, July.
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Jack Dykinga / Aurora Borealis reflected in Polar Lake, conifers silhouetted at edge. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
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Nick Garbutt / Jaguar (Panthera onca palustris), male hunting amongst tree roots at edge of the Cuiaba River. Three Brothers, Porto Jofre, northern Pantanal, Mato Grosso, Brazil. August 2017.
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Guy Edwardes / Lobster pots at edge of Loch Duich, Eilean Donan Castle in background. Highlands, Scotland, UK. January 2014.
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Guy Edwardes / Boat at edge of Loch of Spiggie, Shetland, Shetland Isles, Scotland, UK. July 2014.
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Andy Rouse / Polar bear (Ursus maritimus) standing on pack ice at the water's edge, Svalbard, Norway. June.
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Ross Hoddinott / Catbells and the edge of Derwentwater bathed in early morning light and autumnal colours, Latrigg, Keswick, The Lake District, Cumbria, UK. November 2016.