Michael D. Kern / Kaleidoscope pattern formed from picture of Johnstons Chameleon (Chamaeleo johnstoni) face and legs. Restricted for Editorial use until December 2015

Yukihiro Fukuda / Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan.

Gerry Ellis / Chimpanzee (Pan troglodytes) orphans, Larry investigating Daphne's tongue, Ape Action Africa, Mefou Primate Sanctuary, Cameroon

Juergen & Christine Sohns / African Lion (Panthera leo) male drinking at waterhole, Tswalu Game Reserve, South Africa

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty-three year old female, named Gremlin, scratching face, Gombe National Park, Tanzania

Fiona Rogers / Eastern Chimpanzee (Pan troglodytes schweinfurthii) four year old infant male, named Gizmo, Gombe National Park, Tanzania

Heike Odermatt / Walrus (Odobenus rosmarus), Hinlopen Strait, Svalbard, Norway

Gabriel Rojo / Guanaco (Lama guanicoe) female and newborn baby, nose to nose in grassland. La Pampa Province, Argentina.

Andy Rouse / Bengal tiger (Panthera tigris) male, sub-adult aged three years, face portrait. Ranthambhore National Park, India.

Gary Bell/ Oceanwide / Scarlet Cleaner Shrimp (Lysmata amboinensis) cleaning mouth of Tomato Grouper (Cephalopholis sonnerati), Great Barrier Reef, Australia

Gary Bell/ Oceanwide / Scarlet Cleaner Shrimp (Lysmata amboinensis) cleaning mouth of Tomato Grouper (Cephalopholis sonnerati), Great Barrier Reef, Australia

Gary Bell/ Oceanwide / Whitemouth Moray (Gymnothorax meleagris) being cleaned by Cleaner Wrasse (Labroides sp) juvenile, Christmas Island, Australia

Gary Bell/ Oceanwide / Scarlet Cleaner Shrimp (Lysmata amboinensis) cleaning Yellow-edged Moray (Gymnothorax flavimarginatus), Tulamben, Bali, Indonesia

Gary Bell/ Oceanwide / Scarlet Cleaner Shrimp (Lysmata amboinensis) cleaning Honeycomb Moray Eel (Gymnothorax favagineus), Tulamben, Bali, Indonesia

Gary Bell/ Oceanwide / Variegated Jawfish (Opistognathus solorensis) male brooding eggs in mouth, Anilao, Philippines

Gary Bell/ Oceanwide / Variegated Jawfish (Opistognathus solorensis) male brooding eggs in mouth, Anilao, Philippines

Gary Bell/ Oceanwide / Scarlet Cleaner Shrimp (Lysmata amboinensis) cleaning mouth of Goldbelly Cardinalfish (Apogon apogonides), Tulamben, Bali, Indonesia

Suzi Eszterhas / Leopard (Panthera pardus) mother grooming five-week-old cub, Jao Reserve, Botswana

Neil Aldridge / White rhinoceros (Ceratotherium simum) male and female, face to face, Mosi Oa Tunya National Park, Zambia.

Marie Read / California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA

Marie Read / Great blue heron (Ardea herodias) in breeding plumage, head and shoulders showing plumes on crown and neck, Montezuma National Wildlife Refuge, New York, USA, May.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults brooding chicks. One chick exhibiting aggression towards other. Atka Bay, Antarctica. September.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults with chicks aged six to eight weeks. Atka Bay, Antarctica. September.

Thomas Marent / Buckley's Slender-legged Treefrog (Osteocephalus buckleyi), Putumayo, Colombia

Hermann Brehm / Snow Leopard (Panthera uncia) climbing steep rock face, Mongolia

Suzi Eszterhas / Komodo Dragon (Varanus komodoensis) pair flicking tongues, Komodo National Park, Indonesia

Marion Vollborn/ BIA / Cheetah (Acinonyx jubatus) cub, Castile-La Mancha, Spain

Anup Shah / Olive Baboon (Papio anubis) curious male investigating camera, Masai Mara, Kenya

Staffan Widstrand / Wild Wonders of China / Portrait of a Golden snub-nosed monkey (Rhinopithecus roxellana) full frame of the face, Foping Nature Reserve, Shaanxi, China. Endangered species

David Fleetham / Jaws and teeth of Stout Moray (Gymnothorax eurostus). Hawaii.

David Fleetham / Marine iguana (Amblyrhynchus cristatus) emerging from ocean into Mangroves (Rhizophoraceae). Santa Cruz Island, Galapagos, Ecuador.

David Fleetham / Mouth of the worlds largest fish, the Whale shark (Rhincodon typus), Donsol, Philippines.

David Fleetham / Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.

David Fleetham / White mouthed moray eel (Gymnothorax meleagris), Hawaii.

David Fleetham / Garibaldi (Hypsypops rubicundus) the state fish of California, in forest of giant kelp, (Macrocystis pyrifera) Catalina Island, California, USA.

David Fleetham / Gammarid isopods (Cyproidea sp) on tunicate (Tunicata). Komodo, Indonesia.

David Fleetham / Gammarid isopod (Cyproidea sp) on Tunicate (Tunicata). Komodo, Indonesia.

David Fleetham / Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean.

Sebastian Kennerknecht / Leopard (Panthera pardus) female feeding on sub-adult male Impala (Aepyceros melampus) prey in tree, Greater Makalali Private Game Reserve, South Africa

Sebastian Kennerknecht / American Bison (Bison bison) female cleaning nose with tongue, Yellowstone National Park, Wyoming

Radomir Jakubowski / Red fox (Vulpes vulpes) face portrait in snow, Grand Teton National Park, Wyoming, USA, February.

Lucas Bustamante / Female African lion (Panthera leo) face portrait, Etosha National Park, Harare Province, Namibia.

Claudio Contreras / Young girl face to face with Peninsular Pronghorn Antelope (Antilocapra americana peninsularis) fawn at Peninsular Pronghorn recovery project, El Vizcaino Desert Biosphere Reserve, Baja California Pen...

Tim Fitzharris / Sumatran Orangutan (Pongo abelii) male, native to Sumatra

Paul Bertner / Fly with compound eyes, Khao Yai National Park, Thailand

Ingo Arndt / Head portrait of Eastern grey kangaroo {Macropus giganticus} Australia

Philippe Clement / Juvenile European brown bear cub (Ursus arctos arctos) walking in deep snow in winter, Bavarian Forest National Park, Germany, January. Captive.

Philippe Clement / Group of Alpine ibex (Capra ibex) climbing a cliff face, with others fighting above them, Gran Paradiso National Park, Graian Alps, Italy, June.

Philippe Clement / Three male Alpine ibex (Capra ibex) climbing up a steep mountain rock face, one pushes another one away, Gran Paradiso National Park, Graian Alps, Italy, June.

Philippe Clement / Two male Alpine ibex (Capra ibex) fighting on a steep mountain rock face, Gran Paradiso National Park, Graian Alps, Italy, June.

Steve Downer / Newly hatched slug (Gastropoda) approaching a water droplet on a leaf before bursting it and retracting its eyestalks, Birmingham, England, UK. Controlled conditions.

Steve Downer / Banded snail (Cepaea nemoralis) on rear of a backlit rose leaf, controlled conditions.

Silke Arndt / Western rattlesnake (Crotalus viridis viridis), threat display, South Dakota, USA, September.

Sue Daly / Close up of the face of a Brill (Scophthalmus rhombus), swims out of frame, Sark, British Channel Islands, UK, August.

Sue Daly / Close up of a Plaice (Pleuronectes platessa), swims away, Sark, British Channel Islands, UK, June.

John Waters / Panning shot of farmland along southern face of the Mendip Hills, Somerset, England, UK, August 2014

Ingo Arndt / Baya weaver (Ploceus philippinus) subadult birds practising building 'play nest', Singapore.

Ingo Arndt / King Penguin (Aptenodytes patagonicus) dispute on beach, Volunteer Point, Falkland Islands, February

Ingo Arndt / Yellow-footed rock wallaby (Petrogale xanthopus) Australia, December

Ingo Arndt / Aboriginal Rock Art / depicting human figure / Anbangbang Gallery / Nourlangie Range / Kakadu National Park, Northern Territory, Australia

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Tata' and her unnamed baby aged 2-3 months portrait. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. June 2010. Reh...

Ingo Arndt / Mount Rushmore National Memorial, South Dakota, USA, September 2009

Ingo Arndt / Seabird colony nesting on cliff face, Lange Anna, Helgoland, Germany

Ingo Arndt / Close-up of Owl butterfly (Caligo memnon) face, Central America (Captive)

Ingo Arndt / Red bald headed uakari {Cacajao calvus ucayalii} Rio Yavari, Amazonia, Peru FOR SALE IN UK ONLY

Ingo Arndt / Close up of face of Red bald headed uakari {Cacajao calvus ucayalii} Rio Yavari, Amazonia, Peru FOR SALE IN UK ONLY

Ingo Arndt / Mountain gorilla portrait (Gorilla gorilla berengeii) Parc des Volcans, Rwanda

Ingo Arndt / African wild dog portrait {Lycaon pictus} Namibia. Taken under controlled conditions.

Ingo Arndt / Orang utan male portrait, wild {Pongo pygmaeus} Tanjung Puting National Park, Central Borneo

Ingo Arndt / Cheetah head portrait {Acinonyx jubatus} Namibia

Francois Savigny / Spectacled caiman head on {Caiman sclerops} Llanos, Hato El Frio, Venezuela

Francois Savigny / Male lion washing paw {Panthera leo} Southern Africa

Francois Savigny / Snow leopard portrait {Panthera uncia} captive

Francois Savigny / Snow leopard face portrait {Panthera uncia} captive

Francois Savigny / Snow leopard resting face portrait {Panthera uncia} captive

Francois Savigny / Snow leopard face portrait {Panthera uncia} captive

Ingo Arndt / Fat dormouse in flowering tree {Glis glis} Germany

Ingo Arndt / Japanese macaque sitting portrait {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaque baby portrait {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaque baby {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaque being groomed in hot thermal spring {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaque chewing bark {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaques with young in hot spring {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaques grooming {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaque head portrait {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaque warming in hot thermal spring {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Japanese macaque warming in hot thermal spring {Macaca fuscata} Joshin-etsu NP, Japan

Ingo Arndt / Ornate box turtle retracted into shell {Terrapene ornata} USA

Ingo Arndt / Tiger beetle close up of mouthparts {Cicindela hydriba} Hessen, Germany

Ingo Arndt / Garden spider close-up head portrait {Araneus diadematus} Germany

Ingo Arndt / White-handed gibbon portrait {Hylobates lar} Khao Yai NP, Thailand

Ingo Arndt / Hedgehog eating earthworm {Erinaceus europaeus} Germany

Ingo Arndt / African chameleon {Chamaeleo africanus} portrait, captive, from north-west Africa

Ingo Arndt / Natterjack toad {Bufo calamita} Germany

Francois Savigny / Spectacled caiman head close-up {Caiman crocodilus} Llanos, Venezuela Hato el Frio

Francois Savigny / Tiger (Panthera tigris) in undergrowth. Khana NP, Madhya Pradesh, India