Danny Green / Reindeer (Rangifer tarandus), two play fighting in snow. Svalbard, Norway, April.
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Danny Green / Polar bear (Ursus maritimus) female and cubs play fighting. Svalbard, Norway, March 2018.
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Ann & Steve Toon / African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana.
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Alex Mustard / Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...
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Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
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Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.
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Paul Hobson / Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.
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Axel Gomille / Wolf (Canis lupus), young wolves play fighting in meadow, Saxony-Anhalt, Germany.
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Giri Cavale / Melanistic leopard / Black panther (Panthera pardus) on territorial patrol on track, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India.
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Agustin Esmoris / Southern Elephant Seal (Mirounga leonina) male calling and bleeding after fight, Puerto Madryn, Argentina
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Dietmar Nill / Two European hares (Lepus europaeus) boxing in a snow covered field, Germany, March.
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SCOTLAND: The Big Picture / European river otter (Lutra lutra) cubs play fighting, Shetland, Scotland, UK, May.