Benjamin Barthelemy,Benjamin Barthelemy / Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June

Bence Mate / European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition.

Diane McAllister / Polar bears (Ursus maritimus) fighting in water. Beaufort Sea , Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Nature's Best Photography Competition 2019.

Chase Dekker / Lesser Flamingo (Phoenicopterus minor) pair fighting, Amboseli National Park, Kenya

Danny Green / Reindeer (Rangifer tarandus), two play fighting in snow. Svalbard, Norway, April.

Danny Green / Polar bear (Ursus maritimus) female and cubs play fighting. Svalbard, Norway, March 2018.

Ann & Steve Toon / African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana.

David Tipling / Common European adder (Vipera berus) two males fighting over a female. Holt, North Norfolk, England, UK. April

David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April

David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.

Gary Bell/ Oceanwide / Australian Giant Cuttlefish (Sepia apama) males fighting over female, Whyalla, South Australia, Australia

Gary Bell/ Oceanwide / Australian Giant Cuttlefish (Sepia apama) males fighting over female, Whyalla, South Australia, Australia

Alex Mustard / Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...

Tony Wu / View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.

Tony Wu / Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tony Wu / Zoarchias major eelpouts. (Zoarchias major) The orange individual is female, the one in the hole is male. The female has approached the male, curled around him, attempting to dislodge the male from th...

Tony Wu / Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tui De Roy / Marine iguanas (Amblyrhynchus cristatus) fighting, Punta Vicente Roca, Isabela Island, Galapagos

John Chan / Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...

Wim van den Heever / King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands. 2018 Wildlife Photographer of the Year LUMIX Peo...

Bence Mate / African wild dogs (Lycaon pictus) juveniles playing with the leg of an impala, trying to drag it in three different directions. Mkuze, South Africa. Commended in Glanzlichter photo competition 2020.

Terry Whittaker / Red foxes (Vulpes vulpes) fighting in sand dune habitat, Holland, Netherlands. November.

Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA

Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.

David Pattyn / Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May

Matthew Maran / Red Fox (Vulpes Vulpes) cubs play fighting North London, England, UK, June.

Alan Murphy/ BIA / Northern Caracara (Caracara cheriway) pair fighting, Texas

Alan Murphy/ BIA / Baltimore Oriole (Icterus galbula) males fighting, Texas

Alan Murphy/ BIA / Baltimore Oriole (Icterus galbula) males fighting, Texas

Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.

Markus Varesvuo / Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October.

George Sanker / Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.

Staffan Widstrand / Wild Wonders of China / Pere David's deer / Milu (Elaphurus davidianus), two stags testing their strength in Hubei Tian'ezhou Milu National Nature Reserve, Shishou, Hubei, China

Paul Hobson / Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.

David Perpinan / Group of Griffon vultures (Gyps fulvus) fighting to feed on a dead lamb. Cuenca, Spain, August.

Axel Gomille / Wolf (Canis lupus), young wolves play fighting in meadow, Saxony-Anhalt, Germany.

Andy Rouse / White-necked Jacobin hummingbirds (Florisuga mellivora) fighting, Tobago Small repro only

Ingo Arndt / Guanaco (Lama guanicoe) males fighting, Torres del Paine National Park, Patagonia, Chile

Kerstin Hinze / Two Black-bellied hamsters (Cricetus cricetus) fighting, Vienna, Austria, October.

Tui De Roy / San Cristobal giant tortoises (Chelonoidis chatamensis), fighting, Galapaguera, San Cristobal Island, Galapagos

Philippe Clement / Two Lynx (Lynx lynx) kittens play fighting in front of den entrance, Bavarian Forest National Park, Germany, October. Captive.

Richard Du Toit / Impala (Aepyceros melampus) males fighting, Mkhuze Game Reserve, KwaZulu-Natal, South Africa

Remi Masson / Male Common toads (Bufo bufo) fighting underwater during mating season, with a pair mating Ain, Rhone-Alps, France, March.

Dave Bevan / Slow motion clip of two pairs of Jackdaws (Corvus monedula) squabbling over nest site, Worcestershire, England, UK, May.

Alan Murphy/ BIA / Northern Cardinal (Cardinalis cardinalis) male and Pyrrhuloxia (Cardinalis sinuatus) male fighting, Texas

Sebastian Kennerknecht / African Elephant (Loxodonta africana) bulls fighting, Kruger National Park, South Africa

Sergey Gorshkov / Northern goshawk (Accipiter gentilis) flying with squirrel prey, Finland . March

Sergey Gorshkov / Northern goshawks (Accipiter gentilis) fighting over squirrel carcass, Finland, March.

Sergey Gorshkov / Black Grouse (Tetrao tetrix) males fighting at lek in the snow, Tver, Russia. April

Mark Raycroft / Elk (Cervus elaphus) bulls fighting, North America

Alan Murphy/ BIA / Great Kiskadee (Pitangus sulphuratus) and Green Jay (Cyanocorax yncas) fighting, Texas

Alan Murphy/ BIA / Sharp-tailed Grouse (Tympanuchus phasianellus) males fighting at lek in snow, Minnesota

Alan Murphy/ BIA / Sharp-tailed Grouse (Tympanuchus phasianellus) males fighting at lek in snow, Minnesota

Donald M. Jones / Elk (Cervus elaphus) bulls fighting, western Canada

Dave Bevan / Juvenile male Wild boar (Sus scrofa) fighting to establish seniority, Gloucestershire, England, UK, December.

Philippe Clement / Red deer (Cervus elaphus) stags fighting, Scotland, UK, May.

Erlend Haarberg / Mountain hares (Lepus timidus) fighting in snow at night, Vauldalen, Norway. Joint overall winner of the GDT European Wildlife Photographer of the Year 2017. Highly commended in the Wildlife Photo...

Brian Bevan / Brown hare (Lepus europaeus) boxing, Bedfordshire, England, UK, March.

Dave Bevan / Two male Grey seals (Halichoerus grypus) fighting in surf, with a female and juvenile nearby, Norfolk, England, UK. December.

Francisco Marquez / Two male Iberian wild goats (Capra pyrenaica) fighting, Sierra de Gredos, Spain, November.

Sandesh Kadur / Two Bengal monitors (Varanus bengalensis) fighting, Ranthambore National Park, Rajasthan, India. 2016.

Philippe Clement / Group of Alpine ibex (Capra ibex) climbing a cliff face, with others fighting above them, Gran Paradiso National Park, Graian Alps, Italy, June.

Philippe Clement / Two male Alpine ibex (Capra ibex) fighting on a steep mountain rock face, Gran Paradiso National Park, Graian Alps, Italy, June.

Frederique Olivier / Chinstrap penguin (Pygoscelis antarcticus) adding a stick to its nest and fighting with a neighbouring pair, Aitcho Island, South Shetland Islands.

Sandesh Kadur / Two male King cobras (Ophiophagus hannah) fighting, Agumbe, Karnartaka, India.

Ammonite / Male Yellow dung flies (Scathophaga stercoraria) fighting over a female on a cowpat, UK.

Ammonite / Male Yellow dung flies (Scathophaga stercoraria) fighting over a female on a cowpat, UK.

Dietmar Nill / Two European hares (Lepus europaeus) boxing in a snow covered field, Germany, March.

Konrad Wothe / Showreel of Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.

Ingo Arndt / Gelada baboon males fighting, Simien Mt NP, Ethiopia {Theropithecus gelada}

Ingo Arndt / Eastern grey kangaroos sparring {Macropus giganteus} Australia

Ingo Arndt / Two Northern elephant seal (Mirounga angustirostris) males fighting for dominance, California, USA

Ingo Arndt / Jackson's chameleons fighting {Chamaeleo jacksonii} Kenya

Ingo Arndt / Gelada baboon {Theropithecus gelada} juveniles playing on rocks, Simien Mountains NP, Ethiopia

Konrad Wothe / Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.

Thomas Hinsche/ BIA / Common Buzzard (Buteo buteo) pair fighting, Saxony-Anhalt, Germany

Thomas Hinsche/ BIA / Common Buzzard (Buteo buteo) pair fighting, Saxony-Anhalt, Germany

SCOTLAND: The Big Picture / Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April.

SCOTLAND: The Big Picture / Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April.

SCOTLAND: The Big Picture / European river otter (Lutra lutra) cubs play fighting, Shetland, Scotland, UK, May.

Bence Mate / African wild dogs (Lycaon pictus) play fighting, Mkuze, South Africa

Ingo Arndt / Masai giraffe (Giraffa camelopardalis tippelskirchi) males fighting, Masai Mara National Reserve, Kenya. Sequence 3 of 3

Carol Walker / Two wild Mustang stallions fighting in the White Mountain Herd Area, Wyoming, USA. August.

Sergey Gorshkov / Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa.

Jordi Chias / Two male Dusky groupers (Epinephelus marginatus) fighting for territory during mating season at 'baixa do meros', a small pinnacle at Formigas islets Protected area, near Santa Maria Island, Azores. F...

Danny Green / Coastal brown bears (Ursus arctos) fighting, Lake Clarke National Park, Alaska, September

Will Burrard-Lucas / Aardvark (Orycteropus afer) fighting in the dark. Makgadikgadi Pans, Botswana. January

Yashpal Rathore / Asiatic elephant (Elephas maximus) young female play fighting. Jim Corbett National Park, India. 2014

Bence Mate / Great egrets (Ardea alba) fighting with necks intertwined, Lake Csaj, Kiskunsagi National Park, Hungary. January.

Bence Mate / Juvenile White tailed sea eagles (Haliaeetus albicilla) fighting for fish in mid air, Lake Csaj, Kiskunsagi National Park, Hungary. February.

Bence Mate / Two Buzzards (Buteo buteo) fighting, Lake Csaj, Kiskunsagi National Park, Hungary. February.

Bence Mate / Great white egret (Ardea alba) fighting with Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Hungary, January.

Bence Mate / Caspian gulls (Larus cachinnans) juveniles fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. March.

Bence Mate / Blackbird (Turdus merula) fighting, splashing water, Pusztaszer, Kiskunsagi National Park, Hungary, April.