Ismaele Tortella / White tailed eagle (Haliaeetus albicilla) catching fish, Lofoten, Norway, August.

Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Hermann Brehm / Cuckoo (Cuculus canorus) in flight, Germany, April. May

Sylvain Cordier / Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site, An...

Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.

Nick Garbutt / Black-browed albatross (Thalassarche melanophris) flying over South Atlantic. Cooper Bay, South Georgia. November.

Shane Gross / Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean.

Guy Edwardes / Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.

Ismaele Tortella / White tailed eagle (Haliaeetus albicilla) flying, Lofoten, Norway, August.

Morley Read / Slow motion clip of a large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador. (non-ex)

Morley Read / Twistneck turtle (Platemys platycephala) resting, with butterfly landing on its head, Rio Yasuni, Orellana Province, Ecuador. (non-ex)

Espen Bergersen / Northern fulmar (Fulmarus glacialis) in front of the Seljalandsfoss waterfall, Iceland. May

Espen Bergersen / Herring gull (Larus argentatus) flying over the enormous black beach close to Dyrhoaley, Iceland, May 2016.

Hermann Brehm / Cuckoo (Cuculus canorus) in flight, Germany. April.

Doug Gimesy / Grey-headed flying-foxes (Pteropus poliocephalus) hang in a tree with one coming in to land. Yarra Bend Park. Fairfield, Victoria, Australia. November.

Doug Gimesy / Grey-headed flying-foxes (Pteropus poliocephalus) hang in a tree with one coming in to land. Yarra Bend Park. Kew, Victoria, Australia. November.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) Yarra Bend Park, Kew, Victoria, Australia. November.

Doug Gimesy / Grey-headed flying-foxes (Pteropus poliocephalus) hang from a branch during a light summer rain shower. Yarra Bend Park, Kew, Victoria, Australia. November.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) flying during a very light rain shower with her baby / pup hanging on. Yarra Bend Park, Kew, Victoria, Australia. November.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) flying during a very light rain shower. Yarra Bend Park, Kew, Victoria, Australia. November.

Doug Gimesy / Portrait of a Grey-headed flying-fox (Pteropus poliocephalus) hanging from a branch. Yarra Bend Park, Kew, Victoria, Australia. October.

Doug Gimesy / Portrait of a Grey-headed Flying-fox (Pteropus poliocephalus) hanging from a branch. Yarra Bend Park, Kew, Victoria, Australia. October.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) that has just given birth, eats the placenta. Yarra Bend Park, Kew, Victoria, Australia. October.

Doug Gimesy / Grey-headed Flying-fox (Pteropus poliocephalus) that has just given birth, eats the placenta. Yarra Bend Park, Kew, Victoria, Australia. October.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) female that has just given birth licking clean her new born. Placenta visible. Yarra Bend Park, Kew, Victoria, Australia. October.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) flying over the Yarra River, Melbourne. Watched on by wildlife rescuer Louise Bonomi. Fairfield, Victoria, Australia. March.

Doug Gimesy / Parks Victoria ranger and Grey-headed Flying-fox Project Officer Stephen Brend, releasing a Grey-headed flying-fox (Pteropus poliocephalus) back to Yarra Bend Park, which was in care. Watched on by wi...

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) flying .??Yarra Bend Park, Kew, Victoria, Australia. ?January.

Doug Gimesy / During an extreme heat-stress event where temperatures exceeded 40°C, Melbourne Zoo veterinarian Sarah Frith gives in-filed triage to a suffering young and Grey-headed Flying-fox (Pteropus polioce...

Doug Gimesy / Bat rescuer, carer and founder of Fly-by-night bat clinic, Tamsyn Hogarth, comforts a young rescued Grey-headed Flying-fox (Pteropus poliocephalus) that has just been rescued during a heat-stress even...

Doug Gimesy / Bat rescuer, carer and founder of Fly-by-night bat clinic, Tamsyn Hogarth, comforts a young rescued Grey-headed Flying-fox (Pteropus poliocephalus) that has just been rescued during a heat-stress even...

Doug Gimesy / During an extreme heat-stress event where temperatures exceeded 40°C, wildlife rescuer and carer Kate Chamberlain gives in-field triage to a suffering Grey-headed Flying-fox (Pteropus poliocephalu...

Doug Gimesy / Portrait of a young pup Grey-headed Flying-fox (Pteropus poliocephalus) with piebaldism (a rare genetic condition, typically present at birth, in which an animal develops unpigmented) hangs from a tre...

Doug Gimesy / Portrait of a young pup Grey-headed Flying-fox (Pteropus poliocephalus) with piebaldism (a rare genetic condition, typically present at birth, in which an animal develops unpigmented) hangs from a tre...

Doug Gimesy / During an extreme heat-stress event where temperatures exceeded 40°C at Melbourne's Yarra Bend Grey-headed Flying-fox colony, Melbourne Zoo veterinarian Sarah Frith (right) gives in-filed triage t...

Doug Gimesy / Volunteer bat rescuer Tracy Cleaves, spraying water t to cool down some Grey-headed Flying-foxes (Pteropus poliocephalus) that have descended from the safety of higher branches, in an attempt to escap...

Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed Flying-fox colony, where temperatures exceeded 40°C, Melbourne Zoo veterinarian Sarah Frith gives in-field triage to a suf...

Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed Flying-fox colony, where temperatures exceeded 40°C, Melbourne Zoo veterinarian Sarah Frith gives in-field triage to a suf...

Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed Flying-fox (Pteropus poliocephalus) colony, where temperatures exceeded 40°C, Grey-headed Flying-foxed (Pteropus poliocepha...

Doug Gimesy / 1554 recovered Grey-headed Fly-fox (Pteropus poliocephalus) carcases collected during a heat-stress at event at Melbourne's Yarra Bend Grey-headed Flying-fox colony - about one third of those estimate...

Doug Gimesy / Portrait of 1554 recovered Grey-headed Fly-fox (Pteropus poliocephalus) carcases collected during a heat-stress at event at Melbourne's Yarra Bend Grey-headed Flying-fox colony - about one third of th...

Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed Flying-fox (Pteropus poliocephalus) colony, where temperatures exceeded 43??in a desperate search for somewhere cooler and les...

Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed Flying-fox (Pteropus poliocephalus) colony, where temperatures exceeded 43°C, in a desperate search for somewhere cooler a...

Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed flying-fox (Pteropus poliocephalus) colony, where temperatures exceeded 43°C, Grey-headed flying-fox (Pteropus poliocepha...

Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed Flying-fox (Pteropus poliocephalus) colony, where temperatures exceeded 43°in a desperate search for somewhere cooler and...

Doug Gimesy / Park Ranger and Grey-headed Flying-fox Project Officer Stephen Brend on his way to get more supplies and coordinate rescue attempts, walks past a wheelbarrow filled with dead Grey-headed Flying-foxes...

Doug Gimesy / Firefighter from Melbourne's Metropolitan Fire Brigade (MBF) try and help cool down the Yarra Bend Grey-headed Flying-fox (Pteropus poliocephalus) colony by spraying water on them. Park Ranger and Gre...

Doug Gimesy / Firefighter from Melbourne's Metropolitan Fire Brigade (MBF) try and help cool down the Yarra Bend Grey-headed Flying-fox (Pteropus poliocephalus) colony by spraying water on them. Park Ranger and...

Doug Gimesy / Firefighters from Melbourne's Metropolitan Fire Brigade (MBF) try and help cool down Melbourne's Yarra Bend Grey-headed flying-fox (Pteropus poliocephalus) colony by spraying water on them. Pa...

Doug Gimesy / Wheelbarrow filled with just some dead Grey-headed flying-foxes (Pteropus poliocephalus) that were collected earlier that day, the result of an extreem heat event at Melbourne's Yarra Bend Grey-headed...

Doug Gimesy / Sprinkler system set up high in the trees at Melbourne's Yarra Bend Grey-headed flying-fox (Pteropus poliocephalus) colony in an attempt to try and keep them cool during an extreme heat event whe...

Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed flying-fox (Pteropus poliocephalus) colony, where temperatures exceeded 40°C, wildlife rescuer and carer Kate Chamber...

Doug Gimesy / Portrait of a Grey-headed flying-fox (Pteropus poliocephalus) hanging from a branch with wings wrapped around body to help keep dry during a summer rain shower. Yarra Bend Park, Kew, Victoria, Austra...

Doug Gimesy / Portrait of a Grey-headed flying-fox (Pteropus poliocephalus) hanging from a branch with wings wrapped around body to help keep dry during a summer rain shower. Yarra Bend Park, Kew, Victoria, Austr...

Doug Gimesy / Portrait of a Grey-headed flying-fox (Pteropus poliocephalus) hanging from a branch with wings wrapped around body to help keep dry during a summer rain shower. Yarra Bend Park, Kew, Victoria, Austra...

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) hanging from a branch. Yarra Bend Park, Kew, Victoria, Australia. December. Cropped.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) in flight flares wings coming in to land. Yarra Bend Park, Kew, Victoria, Australia. December 2019.?

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) in flight flares wings coming in to land, during a light summer rain shower. Yarra Bend Park, Kew, Victoria, Australia. December.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) female in flight with baby/pup hanging on underneath. Yarra Bend Park, Kew, Victoria, Australia. December 2019.

Doug Gimesy / Trees with Grey-headed flying-foxes (Pteropus poliocephalus) hang in them. Yarra Bend Park. Kew, Victoria, Australia. November 2019.

Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) spotting and flaring wings to come in to land on a branch. Yarra Bend Park, Kew, Victoria, Australia. November 2019.

Doug Gimesy / Grey-headed flying-foxes (Pteropus poliocephalus) hang from a branch.?Yarra Bend Park, Kew, Victoria, Australia. November 2019.

Doug Gimesy / Portrait of a male Grey-headed flying-fox (Pteropus poliocephalus) coming in to land. Yarra Bend Park, Kew, Victoria, Australia.? November 2019.

Peter Lewis / Mediterranean gull (Larus melanocephalus) in Summer plumage in flight, Hayling Island, Hampshire, England, UK.

Peter Lewis / Mediterranean gull (Larus melanocephalus) in Summer plumage in flight, Hayling Island, Hampshire, England, UK.

Peter Lewis / Common Cuckoo (Cuculus canorus) wings open landing on post Surrey, England, UK.

Peter Lewis / Common Cuckoo (Cuculus canorus) wings open landing on post Surrey, England, UK.

Peter Lewis / Common Cuckoo (Cuculus canorus) wings open landing on post Surrey, England, UK.

Peter Lewis / Mediterranean gull (Larus melanocephalus) in summer plumage in flight, Hayling Island, Hampshire, England, UK. July.

Bernard Castelein / Barn swallow (Hirundo rustica) two juveniles, one in flight, the other resting on roof. Klein Schietveld, Brasschaat, Belgium. August.

Oscar Dewhurst / Brahminy kite (Haliastur indus) in flight. Goa, India.

Oscar Dewhurst / Brahminy kite (Haliastur indus) in flight over sea, catching fish in coastal waters. Goa, India.

Oscar Dewhurst / Brahminy kite (Haliastur indus) in flight over sea, catching fish in coastal waters. Goa, India.

Oscar Dewhurst / Malabar pied hornbill (Anthracoceros coronatus) male in flight. Goa, India. September.

Hanne & Jens Eriksen / Bridled tern (Onychoprion anaethetus) in flight. Oman, June.

Hanne & Jens Eriksen / Black-billed magpie (Pica hudsonia) in flight. Colorado, USA, August.

Hanne & Jens Eriksen / Swainson's hawk (Buteo swainsoni) in flight. Saskatchewan, Canada, August.

Hanne & Jens Eriksen / Ruddy turnstone (Arenaria interpres) in flight over sea. Oman, November.

Hanne & Jens Eriksen / Pheasant-tailed jacana (Hydrophasianus chirurgus) male in breeding plumage, in flight. Oman, July.

Hanne & Jens Eriksen / Persian shearwater (Puffinus persicus) taking off, running over water. Oman, October.

Hanne & Jens Eriksen / Northern harrier (Circus hudsonius) juvenile in flight. Saskatchewan, Canada, August.

Hanne & Jens Eriksen / Great horned owl (Bubo virginianus) in flight. Saskatchewan, Canada, August.

Hanne & Jens Eriksen / Crab-plover (Dromas ardeola) flying with crab in beak. Oman, June.

Hanne & Jens Eriksen / Crab-plover (Dromas ardeola) three flying, two with crabs in beaks. Oman, June.

David Allemand / Northern fulmar (Fulmarus glacialis) in flight with waves, Arctic Sea. Commended in the Biophoto Competition 2019.

Bence Mate / African elephant (Loxodonta africana) two silhouetted at sunset with goose flying overhead, Mkuze, South Africa. Highly commended in the African Wildlife category of the Nature's Best Photography...

Morley Read / Slow motion clip of a Spiny orb weaver (Micrathena) biting a flying ant caught in its web prior to eating it, Napo Province, Ecuadorian Amazon. (non-ex)

Ann & Steve Toon / Tawny eagle (Aquila rapax) in flight, Zimanga private game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Reed cormorant (Microcarbo africanus) flying from water with fish, Chobe river, Botswana.

Ann & Steve Toon / Cape sugarbird (Promerops cafer), Kirstenbosch National Botanical Garden, Cape Town, South Africa.

Ann & Steve Toon / Martial eagle (Polemaetus bellicosus) juvenile in flight, Zimanga private game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / African fish eagle (Haliaeetus vocifer) carrying nesting material, Chobe river, Botswana.

Roger Powell / Sandwich Tern (Sterna sandvicensis) flying off with a fish in its beak. Warkworth, Northumberland, England, UK, May.

Roger Powell / Peregrine Falcon (Falco peregrinus) in flight. Fairburn RSPB Reserve, Yorkshire, England, UK, August.