Yashpal Rathore / Bengal Tiger (Panthera tigris) foot print / pugmark on wet sand, Tadoba National Park, India

David Woodfall / Long legged flies (Dolichopodidae) on puddle in foot print of Long Horn cattle. Knepp Wildland Project, formerly intensive farmland now turned to conservation and sustainable farming. Horsham, West Su...

Ashley Cooper / Derwent Water at Keswick in the Lake District completely frozen over during the December 2010 big chill, with footsteps from a walker who walked across the lake. December 2010

Ashley Cooper / Footprint in snow, Ambleside, Lake District, Cumbria, England, UK. February 2009

SCOTLAND: The Big Picture / Otter (Lutra lutra) tracks on beach, Sutherland, Scotland, UK, November

Paul D Stewart / Lithograph of the Moody Footmark Quarry in South Hadley, where Pliny Moody discovered the very first fossil tracks in 1802. From Hitchcock, Edward. Ichnology of New England. A Report on the Sandstone...

Laurent Geslin / Foot print of a female European lynx (Lynx lynx) Jura, Switzerland, February.

David Kjaer / Landscape of Martin Down National Nature Reserve with foot path through grassy downland, Hampshire, England, UK, June 2014.

Stephen Belcher / Javan Rhinoceros (Rhinoceros sondaicus) foot print being examined by Rhino Protection Unit Ranger, Ujung Kulon National Park, Indonesia

Stephen Belcher / Sumatran Rhinoceros (Dicerorhinus sumatrensis) plaster cast of juvenile and mature sumatran rhino foot print, Way Kambas National Park, Sumatra, Indonesia

Stephen Belcher / Sumatran Rhinoceros (Dicerorhinus sumatrensis) plaster cast of juvenile foot print, Way Kambas National Park, Sumatra, Indonesia

Niall Benvie / European lynx (Lynx lynx) footprint in mud, with birch leaf, Sumava NP, Bohemia, Czech Republic.