SCOTLAND: The Big Picture / Mountain hare footprints in snow on mountain top in Glen Coe, Lochaber, Scotland, UK.February

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) footprints on sea ice. Atka Bay, Antarctica. August.

Ingo Arndt / Brown / Grizzly Bear (Ursus arctos horribilis) tracks on tidal flats, Alaska, USA

Ingo Arndt / Desert chameleon crossing sand leaving tracks, Namibia {Chamaeleo namaquensis}

Colin Monteath/ Hedgehog House / Spinifex Grass (Spinifex sp) on sand dune with footprints, Simpson Desert, Northern Territory, Australia

Ole Jorgen Liodden / Polar bear (Ursus maritimus) footprints in snow, on cracked ice, Svalbard, Norway, September.

Jabruson / African forest elephant (Loxodonta africana cyclotis) footprint in drying mud, Romani Bai, Odzala-Kokoua National Park, Democratic Republic of Congo.

Ernie Janes / Oystercatcher (Haematopus ostralegus) footprints on sand at low tide, Norfolk, England, UK, November.

Mark Hamblin / 2020VISION / Mountain hare (Lepus timidus) footprints in snow, Creag Meagaidh National Nature Reserve, Scotland, UK, December 2012.

Paul D Stewart / Landscape of the Triassic coastal environment with reconstructions of dinosaurs and marine reptiles, 'Dr. Schubert's Naturgeschichte - Geologie, Mineralreich, Palaontologie'. Published in Stuttgart 18...

Erlend Haarberg / Muskox (Ovibos moschatus) footprints left in snow, Dovrefjell-Sunndalsfjella National Park, Norway, December 2007

Mark Hamblin / 2020VISION / European river otter (Lutra lutra) footprints crossing frozen Loch Lurgainn at dawn, with Sgorr Tuath and Sgorr Deas (Beinn an Eoin) behind, Coigach, Wester Ross, Scotland, UK, December 2010