Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...

Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...

Konrad Wothe / Fritillary butterfly (Argynnis sp) with Scarce copper butterflies (Lycaena virgaureae) Queyras, Alps, France. August.

Nick Upton / Slow motion clip of a Meadow brown butterfly (Maniola jurtina) and a Gatekeeper butterfly (Pyronia tithonus) nectaring on Saw-wort flowers (Serratula tinctoria), with a Silver-washed fritillary (Argyn...

Nick Upton / Close-up of a Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower and flying off, Wiltshire, England, UK, July.

Lorraine Bennery / Knapweed fritillary butterflies (Melitaea phoebe) on Green winged orchid (Anacamptis morio), Grands Causses Regional Natural Park, France, May.

Dave Watts / Dark green fritillary (Argynnis aglaja) on pink clover, south west France.

Jussi Murtosaari / Pallas Fritillary (Argynnis laodice) portrait of female head on, southern Finland, August.

Jussi Murtosaari / Heath Fritillary (Melitaea athalia) in flight, Finland June

David Woodfall / Small Pearl bordered fritillary butterfly (Boloria selene) in grass, Radnorshire Wildlife Trust Nature Reserve, Wales, UK, June.

Sven Zacek / Fritillary butterfly (Melitaea sp.) – either Heath or False Heath – at rest on a grass stem. Estonia, Europe, June.

David Kjaer / Pearl-bordered Fritillary butterfly (Boloria euphrosyne) at rest on Bracken with wings open, Bentley Wood, Wiltshire, England, UK