Alex Mustard / Backlit detail of branching fire coral (Millepora alcicornis) showing the hair like polyps, filled with stinging nematocysts. East End, Grand Cayman, Cayman Islands. Caribbean Sea.

Cyril Ruoso / Atlantic puffin (Fratercula arctica) seen from burrow, Iceland. July.

Gabriel Rojo / Scimitar-billed woodcreeper (Drymornis bridgesii) at nest hole, Calden forest, La Pampa, Argentina.

Espen Bergersen / Kittiwakes (Rissa tridactyla) squabbling on ice. Svalbard, Norway. July

Cheryl-Samantha Owen / Cape gannet (Morus capensis) colony, Bird Island Nature Reserve, Lambert's Bay, West Coast, South Africa.

Pal Hermansen / Willow warbler (Phylloscopus trochilus) feeding young on steering wheel of abandoned car in 'car graveyard' Varmland, Sweden, June

Diane McAllister,Diane McAllister / Japanese Macaque (Macaca fuscata) females gather at one end of the hot springs, Jigokudani, Japan, February

Juan Carlos Munoz / Gelada baboon (Theropithecus gelada) troop, Simien Mountains, Ethiopia, Africa

George Sanker / Black bear (ursus americanus) cub standing on hind legs, Yellowstone National Park, USA, June

Laurie Campbell / Mute swans (Cygnus olor) newly hatched cygnets on pond, Yetholm Loch Scottish Wildlife Trust Reserve, Roxburghshire, Scotland, May

Bence Mate / Head portraits of King vulture (Sarcoramphus papa) right, and Black vulture (Coragyps atratus) left. Santa Rita, Costa Rica. WINNER: Eric Hosking Award portfolio image 6/6 - Wildlife Photographer of t...

David Tipling / Flock of Rooks (Corvus frugilegus) sitting on telephone wires, at pre-roost gathering in winter, Buckenham, Norfolk, England