Mark Carwardine / Great white shark (Carcharodon carcharias) underwater, Guadalupe Island, Mexico (North Pacific)

Phil Savoie / Smeathman's furrow bee (Lasioglossum smeathmanellum), feeding on wild alpine strawberry (Fragaria vesca), Monmouthshire, Wales, UK. December

Oscar Dewhurst / Mute swan (Cygnus olor), two cygnets sheltering under parent's wing. Richmond Park, London, England, UK. April.

Guy Edwardes / Marbled white (Melanagria galathea), two butterflies on Pyramidal orchid (Anacamptis pyramidalis). Badbury Rings, Wimborne Minster, Dorset, England, UK. June.

George Sanker / Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.

George Sanker / Great egret (Ardea alba) diving head into water to catch prey, in evening light. Myakka River State Park, Florida, USA, March. Sequence 1/2.

Nick Upton / Juvenile Goldenrod spider (Misumena vatia) camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, reacting to and catching a fly (Muscidae), Wiltshire, England, UK, June.

Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...

Nick Upton / Male Thick-legged flower beetle (Oedemera nobilis), with a Pyramidal orchid (Anacamptis pyramidalis) pollinium attached to its head, nectaring on Common hogweed flowers (Heracleum sphondylium), Wiltsh...

Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.

Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.

Nigel Cattlin / Immature caterpillar of a codling moth (Cydia pomonell) eating its way through a gallery in a ripening apple fruit, Berkshire, July