Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...

Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...

Nick Upton / Close-up of a Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower and flying off, Wiltshire, England, UK, July.

Nick Upton / Dark green fritillary butterfly (Argynnis aglaja) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.

Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja), Marbled white butterflies (Melanargia galathea) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed...

Nick Upton / Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, joined by a Meadow brown butterfly (Maniola jurtina), Wiltshire, Englan...

Nick Upton / Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshi...

Rod Williams / Crab spider (Misumena vatia) with Silver-washed fritillary (Argynnis paphia) prey, New Cross Gate Cutting Nature Reserve, Lewisham August.

Nick Upton / Slow motion close up of a Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower alongside a Solitary mining bee (Andrena), Wiltshire, England, UK, July.

Nick Upton / Slow motion clip of a Dark green fritillary butterfly (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower and being joined by a female Thick-legged flower beetle (Oedemera no...

Nick Upton / Slow motion close up view of a Dark green fritillary (Argynnis aglaja)landing to feed from a Greater knapweed (Centaurea scabiosa) flower and displacing a Marmalade hoverfly (Episyrphus balteatus), Wi...

Nick Upton / Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower and flying off, Wiltshire, England, UK, July.