Andy Rouse / Bengal tiger (Panthera tigris tigris) female 'T19 Krishna' with 11 month cub, Ranthambore National Park, India.
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Anup Shah / Celebes / Black crested macaque (Macaca nigra) two juveniles approaching with curiosity, one grimacing Tangkoko National Park, Sulawesi, Indonesia.
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Charlie Summers / African lion (Panthero leo) male giving flehmen grimace, Tanzania, East Africa
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Suzi Eszterhas / Vet examining Sumatran orangutan (Pongo abelii) female rescued by Human Orangutan Conflict Response Unit. Rescued from area of clearcut forest and later released into a national park. Aceh Province, S...
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Glitter' aged 14 years grimacing as part of greeting while being protective over new born baby aged 5-6 days. Gombe National Park, Tanzania...
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Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) baby female, nine month old, grimacing, Gombe National Park, Tanzania
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Wim van den Heever / Jaguar (Panthera onca) yawning, Pantanal Brazil.
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Mark Bowler / Brown capuchin monkey (Sapajus macrocephalus). displaying submissive behavior. Captive, Peru.
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Mark Bowler / Brown capuchin monkey (Sapajus macrocephalus). displaying submissive behavior. Captive, Peru.
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Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) twenty three year old female, named Nasa, making a defensive grimace, Gombe National Park, Tanzania
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Fiona Rogers / Japanese Macaque (Macaca fuscata) juvenile making distress expression. Jigokudani Yaen-Koen National Park, Japan, February.
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Fiona Rogers / Japanese Macaque (Macaca fuscata) baring teeth in stressful situation, Jigokudani Monkey Park, Japan
