Ismaele Tortella / Eurasian otter (Lutra lutra) hunting in a shoal of Atlantic herring (Clupea harengus), northern Norway, March

Ismaele Tortella / Northern gannet (Morus bassanus) colony, chick climbing back into nest, with the midnight sun, Norway, July

Ismaele Tortella / Three Rock ptarmigan (Lagopus muta) feeding in winter plumage, city of Tromso in the background, northern Norway, March 2019.

Ismaele Tortella / Black-legged kittiwake (Rissa tridactyla) nesting above a window, Lofoten, Norway, August.

Ismaele Tortella / Reindeer (Rangifer tarandus), Kvaloya island, Tromso, Norway, November.

Morley Read / Leaf cutter ants (Atta sp) carrying leaves and pieces of yellow flowers along a branch, Amazon rainforest, Orellana Province, Ecuador.

Morley Read / Leaf cutter ants (Atta sp) carrying leaves and pieces of yellow flowers along a branch, Amazon rainforest, Orellana Province, Ecuador.

Morley Read / Leaf cutter ants (Atta sp) carrying leaves and pieces of yellow flowers along a branch, Amazon rainforest, Orellana Province, Ecuador.

Morley Read / Column of Leaf cutter ants (Atta sp) carrying leaves over the rainforest floor to their nest, Amazon rainforest, Orellana Province, Ecuador.

Morley Read / Slow motion clip of a large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador.

Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador.

Morley Read / Slow motion clip of a Leaf cutter ant (Atta) carrying a leaf along a branch, Orellana Province, Ecuador.

Morley Read / Millipede (Diplopoda) walking along a branch, Orellana Province, Ecuador.

Morley Read / Millipede (Diplopoda) walking along a branch, Orellana Province, Ecuador.

Morley Read / Drone shot tracking over a field planted with potatoes, Cornwall, England, UK, June 2018.

Morley Read / Slow motion clip of a Mute swan (Cygnus olor) with three cygnets, Cornwall, England, UK, May.

Morley Read / Slow motion clip of a group of male Mallards (Anas platyrhynchos) competing violently to mate with a single female, Cornwall, England, UK, May 2018.

Sylvain Cordier / Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site, An...

Morley Read / Slow motion clip of a queen termite (Isoptera) in chamber, with workers gathering liquid containing pheromones exuded from the queen's rear end, Ecuador.

Morley Read / Queen Termite (Isoptera) tended by workers in nest chamber, Napo Province, Ecuadorian Amazon.

Morley Read / Slow motion clip of Leafcutter ants (Atta) carrying pieces of leaves back to their nest, Amazon rainforest, Napo Province, Ecuador.

Morley Read / Aerial shot of a group of wild ponies (Equus caballus) in paramo landscape at the base of Cotopaxi Volcano, Ecuadorian Andes, 2018.

Chase Dekker / Humpback Whale (Megaptera novaeangliae) pod gulp feeding on Northern Anchovy (Engraulis mordax), Monterey Bay, California

Chase Dekker / Lesser Flamingo (Phoenicopterus minor) wading, Amboseli National Park, Kenya

Chase Dekker / Lesser Flamingo (Phoenicopterus minor) pair fighting, Amboseli National Park, Kenya

Juergen & Christine Sohns / African Wild Dog (Lycaon pictus) group attacking Cape Warthog (Phacochoerus aethiopicus), Tswalu Game Reserve, South Africa

Heike Odermatt / Brunnich's Guillemot (Uria lomvia) nesting colony on cliff, Alkefjellet, Hinlopen Strait, Svalbard, Norway

Danny Green / Polar bear (Ursus maritimus) female resting with cubs on glacier. Svalbard, Norway, July.

Danny Green / Polar bear (Ursus maritimus) female and cubs feeding on Whale carcass. Svalbard, Norway, July.

Danny Green / Polar bear (Ursus maritimus) and two cubs feeding on Whale carcass. Svalbard, Norway, July.

Danny Green / Polar bear (Ursus maritimus) and cubs, standing on sea ice. Svalbard, Norway, July.

Danny Green / Polar bear (Ursus maritimus) and cubs resting on sea ice. Svalbard, Norway, July.

Danny Green / Polar bear (Ursus maritimus) female and cubs, one cub walking up slope of ice in foreground. Svalbard, Norway, July.

Danny Green / Polar bear (Ursus maritimus) and cubs feeding on Whale carcass. Svalbard, Norway, July.

Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.

Danny Green / Polar bear (Ursus maritimus) group on ice, female and cubs feeding on Whale carcass. Svalbard, Norway, June 2018.

Daniel Heuclin / Llama (Lama glama) herd, Laguna colorada. Altiplano, Bolivia.

Eduardo Blanco / Pale burrower mayfly (Ephoron virgo), swarming in the millions on street at night, appearing like snow. Mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August 2019. Finalist i...

Tony Wu / Bowhead whale (Balaena mysticetus), group socialising in shallow water, aerial view. Part of larger group of more than 50 whales congregating, Sea of Okhotsk sub-population, Russia.

Ann & Steve Toon / African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana.

Tony Wu / Humphead parrotfish (Bolbometopon muricatum) shoal spawning in early morning, part of mass aggregation of thousands of fish. Caroline Islands, Palau.

Alex Mustard / Green turtle (Chelonia mydas) resting on sea floor and two juvenile Golden trevally (Gnathanodon speciosus). Marsa El Shouna, Marsa Alam, Egypt.

Alex Mustard / Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt.

Oscar Dewhurst / Mute swan (Cygnus olor) swimming with cygnets in misty lake in morning light. Richmond Park, London, England, UK. May.

Clay Bolt / Blue-banded bee (Amegilla sp) males sleeping on stem. Halmahera, North Maluku, Indonesia.

Felis Images / Asiatic lion (Panthera leo persica), female and cubs in morning light, cub biting mother's leg. Gir National Park, Gujarat, India. Photo Phillip Ross/Felis Images

Tony Wu / Sperm whale (Physeter macrocephalus), ten swimming together in gathering of more than 50 individuals. Whales with flatulence and defecating. Sri Lanka.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled together under the Southern Lights, Aurora australis, Atka Bay, Antarctica. September.

Gary Bell/ Oceanwide / Australian Pelican (Pelecanus conspicillatus) flock, New South Wales, Australia

Gary Bell/ Oceanwide / Grey Nurse Shark (Carcharias taurus), Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Sea Goldie (Pseudanthias squamipinnis) school in coral reef with feather stars, Great Barrier Reef, Queensland, Australia

Gary Bell/ Oceanwide / Australian Giant Cuttlefish (Sepia apama) group gathered for mating, Whyalla, South Australia, Australia

Gary Bell/ Oceanwide / Australian Sea Lion (Neophoca cinerea) group, Hopkins Island, South Australia, Australia

Gary Bell/ Oceanwide / Australian Sea Lion (Neophoca cinerea) group, Hopkins Island, South Australia, Australia

Gary Bell/ Oceanwide / Piling with sponges, tunicates, and ascidians, Yorke Peninsula, South Australia, Australia

Gary Bell/ Oceanwide / Sea Anemone (Corynactis australis) group, Bicheno, Tasmania, Australia

Gary Bell/ Oceanwide / Yellow-banded Snapper (Lutjanus adetii) school, Heron Island, Great Barrier Reef, Queensland, Australia

Gary Bell/ Oceanwide / Black-spot Goatfish (Parupeneus signatus) and Goldlined Seabream (Rhabdosargus sarba) school, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Blacktip Bullseye (Pempheris affinis) school, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Stripey (Microcanthus strigatus) school, Coffs Harbor, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Eastern Pomfred (Schuettea scalaripinnis) school, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Common Lionfish (Pterois volitans) hunting Basslets (Pseudanthias sp), Christmas Island, Australia

Gary Bell/ Oceanwide / Painted Sweetlips (Diagramma pictum) in Slender Cardinalfish (Rhabdamia gracilis) school, Great Barrier Reef, Australia

Gary Bell/ Oceanwide / Painted Sweetlips (Diagramma pictum) in Slender Cardinalfish (Rhabdamia gracilis) school near Double-headed Maori Wrasse (Cheilinus undulatus), Great Barrier Reef, Australia

Gary Bell/ Oceanwide / Double-headed Maori Wrasse (Cheilinus undulatus), Painted Sweetlips (Diagramma pictum), and Slender Cardinalfish (Rhabdamia gracilis) school, Great Barrier Reef, Queensland, Australia

Tony Heald / Elephant (Loxodonta africana) babies playing, Etosha National Park,Namibia.

Bence Mate / Dying Common frogs (Rana temporaria) with their legs removed for food, left to die in their breeding pool surrounded by frogspawn. Covasna, Romania. Highly commended in the Wildlife Photojournalism Ca...

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

Oscar Dewhurst / Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December

Jack Dykinga / Bighorm sheep (Ovis canadensis) males grouped together before the rutting season. Valley of Fire State Park, Nevada, USA, February.

Jordi Chias / Gentoo penguins (Pygoscelis papua) swimming together and porpoising in search of krill, Antarctic Peninsula, Antarctica.

Jordi Chias / Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica.

Jordi Chias / Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica.

Jordi Chias / Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica.

Ingo Arndt / Female Puma (Puma concolor) with two cubs aged around four months, Torres del Paine National Park, Patagonia, Chile, July.

Ingo Arndt / Aerial shot over a herd of Guanacos (Lama guanicoe), Torres del Paine National Park, Patagonia, Chile, April.

Alex Mustard / Killer whales (Orcinus orca) at the surface in an Arctic fjord in Spildra, northern Norway, with sea birds. Arctic Ocean

Alex Mustard / School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean

Alex Mustard / Shoal of three Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.

Alex Mustard / Pod of False killer whales (Pseudorca crassidens) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka.

Alex Mustard / Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Lucas Bustamante / Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador

Terry Whittaker / Man sorting seabird eggs, including those of Common guillemots (Uria aalge) into egg boxes, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.

Franco Banfi / Pod of sperm whale socializing, (Physeter macrocephalus), Dominica, Caribbean Sea, Atlantic Ocean.

Jack Dykinga / Bald eagle (Haliaeetus leucocephalus) perched as Snow geese (Chen caerulescens) fly by, Bosque del Apache National Wildlife Refuge, New Mexico, USA, January.

Jack Dykinga / Snow goose (Chen caerulescens) landing into a tight packed flock on lake, Bosque del Apache National Wildlife Refuge, New Mexico, USA. January.

Ernie Janes / Brent geese (Branta bernicla) flying with moon, Titchwell, Norfolk, UK, December

Suzi Eszterhas / African Elephant (Loxodonta africana) herd drinking at waterhole, Mashatu Game Reserve, Botswana

Suzi Eszterhas / African Elephant (Loxodonta africana) herd drinking at waterhole, Mashatu Game Reserve, Botswana

John Chan / California brown pelicans (Pelecanus occidentalis californicus) roosting on a mudflat, with a flock of Elegant terns (Thalasseus elegans) nearby, Southern California, USA May/2019

John Chan / Flock of Western sandpipers (Calidris mauri) taking flight from salt marsh, Southern California, USA.

David Woodfall / Lesser horseshoe bats (Rhinolophus hipposideros) in magnesium mine, Shropshire, England, UK, April.

John Shaw / King penguins (Aptenodytes patagonicus) walking in line on a windy beach. Sanders Island, Falklands. November.

John Shaw / Chestnut-eared aracari (Pteroglossus castanotis) on cat-gut vine. Pantanal, Brazil, August.

John Shaw / Roseate spoonbill (Platalea ajaja) at its nesting colony site, in early morning light. St. Johns Management Area, Florida, USA, April.