Ismaele Tortella / Eurasian otter (Lutra lutra) hunting in a shoal of Atlantic herring (Clupea harengus), northern Norway, March

Ismaele Tortella / Northern gannet (Morus bassanus) colony, chick climbing back into nest, with the midnight sun, Norway, July

Ismaele Tortella / Reindeer (Rangifer tarandus), Kvaloya island, Tromso, Norway, November.

Morley Read / Slow motion clip of a group of male Mallards (Anas platyrhynchos) competing violently to mate with a single female, Cornwall, England, UK, May 2018.

Sylvain Cordier / Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site, An...

Morley Read / Aerial shot of a group of wild ponies (Equus caballus) in paramo landscape at the base of Cotopaxi Volcano, Ecuadorian Andes, 2018.

Chase Dekker / Humpback Whale (Megaptera novaeangliae) pod gulp feeding on Northern Anchovy (Engraulis mordax), Monterey Bay, California

Chase Dekker / Lesser Flamingo (Phoenicopterus minor) wading, Amboseli National Park, Kenya

Chase Dekker / Lesser Flamingo (Phoenicopterus minor) pair fighting, Amboseli National Park, Kenya

Juergen & Christine Sohns / African Wild Dog (Lycaon pictus) group attacking Cape Warthog (Phacochoerus aethiopicus), Tswalu Game Reserve, South Africa

Heike Odermatt / Brunnich's Guillemot (Uria lomvia) nesting colony on cliff, Alkefjellet, Hinlopen Strait, Svalbard, Norway

Danny Green / Polar bear (Ursus maritimus) female and cubs, one cub walking up slope of ice in foreground. Svalbard, Norway, July.

Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.

Danny Green / Polar bear (Ursus maritimus) group on ice, female and cubs feeding on Whale carcass. Svalbard, Norway, June 2018.

Daniel Heuclin / Llama (Lama glama) herd, Laguna colorada. Altiplano, Bolivia.

Tony Wu / Bowhead whale (Balaena mysticetus), group socialising in shallow water, aerial view. Part of larger group of more than 50 whales congregating, Sea of Okhotsk sub-population, Russia.

Ann & Steve Toon / African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana.

Tony Wu / Humphead parrotfish (Bolbometopon muricatum) shoal spawning in early morning, part of mass aggregation of thousands of fish. Caroline Islands, Palau.

Alex Mustard / Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt.

Tony Wu / Sperm whale (Physeter macrocephalus), ten swimming together in gathering of more than 50 individuals. Whales with flatulence and defecating. Sri Lanka.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled together under the Southern Lights, Aurora australis, Atka Bay, Antarctica. September.

Gary Bell/ Oceanwide / Australian Pelican (Pelecanus conspicillatus) flock, New South Wales, Australia

Gary Bell/ Oceanwide / Grey Nurse Shark (Carcharias taurus), Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Sea Goldie (Pseudanthias squamipinnis) school in coral reef with feather stars, Great Barrier Reef, Queensland, Australia

Gary Bell/ Oceanwide / Australian Giant Cuttlefish (Sepia apama) group gathered for mating, Whyalla, South Australia, Australia

Gary Bell/ Oceanwide / Piling with sponges, tunicates, and ascidians, Yorke Peninsula, South Australia, Australia

Gary Bell/ Oceanwide / Sea Anemone (Corynactis australis) group, Bicheno, Tasmania, Australia

Gary Bell/ Oceanwide / Yellow-banded Snapper (Lutjanus adetii) school, Heron Island, Great Barrier Reef, Queensland, Australia

Gary Bell/ Oceanwide / Black-spot Goatfish (Parupeneus signatus) and Goldlined Seabream (Rhabdosargus sarba) school, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Blacktip Bullseye (Pempheris affinis) school, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Stripey (Microcanthus strigatus) school, Coffs Harbor, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Eastern Pomfred (Schuettea scalaripinnis) school, Solitary Islands Marine Park, New South Wales, Australia

Gary Bell/ Oceanwide / Common Lionfish (Pterois volitans) hunting Basslets (Pseudanthias sp), Christmas Island, Australia

Gary Bell/ Oceanwide / Painted Sweetlips (Diagramma pictum) in Slender Cardinalfish (Rhabdamia gracilis) school, Great Barrier Reef, Australia

Gary Bell/ Oceanwide / Painted Sweetlips (Diagramma pictum) in Slender Cardinalfish (Rhabdamia gracilis) school near Double-headed Maori Wrasse (Cheilinus undulatus), Great Barrier Reef, Australia

Gary Bell/ Oceanwide / Double-headed Maori Wrasse (Cheilinus undulatus), Painted Sweetlips (Diagramma pictum), and Slender Cardinalfish (Rhabdamia gracilis) school, Great Barrier Reef, Queensland, Australia

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

Oscar Dewhurst / Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December

Jack Dykinga / Bighorm sheep (Ovis canadensis) males grouped together before the rutting season. Valley of Fire State Park, Nevada, USA, February.

Jordi Chias / Gentoo penguins (Pygoscelis papua) swimming together and porpoising in search of krill, Antarctic Peninsula, Antarctica.

Jordi Chias / Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica.

Ingo Arndt / Aerial shot over a herd of Guanacos (Lama guanicoe), Torres del Paine National Park, Patagonia, Chile, April.

Alex Mustard / Killer whales (Orcinus orca) at the surface in an Arctic fjord in Spildra, northern Norway, with sea birds. Arctic Ocean

Alex Mustard / School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean

Alex Mustard / Pod of False killer whales (Pseudorca crassidens) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka.

Alex Mustard / Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean.

Franco Banfi / Pod of sperm whale socializing, (Physeter macrocephalus), Dominica, Caribbean Sea, Atlantic Ocean.

Jack Dykinga / Bald eagle (Haliaeetus leucocephalus) perched as Snow geese (Chen caerulescens) fly by, Bosque del Apache National Wildlife Refuge, New Mexico, USA, January.

Jack Dykinga / Snow goose (Chen caerulescens) landing into a tight packed flock on lake, Bosque del Apache National Wildlife Refuge, New Mexico, USA. January.

Ernie Janes / Brent geese (Branta bernicla) flying with moon, Titchwell, Norfolk, UK, December

Suzi Eszterhas / African Elephant (Loxodonta africana) herd drinking at waterhole, Mashatu Game Reserve, Botswana

Suzi Eszterhas / African Elephant (Loxodonta africana) herd drinking at waterhole, Mashatu Game Reserve, Botswana

John Chan / California brown pelicans (Pelecanus occidentalis californicus) roosting on a mudflat, with a flock of Elegant terns (Thalasseus elegans) nearby, Southern California, USA May/2019

John Chan / Flock of Western sandpipers (Calidris mauri) taking flight from salt marsh, Southern California, USA.

John Shaw / Roseate spoonbill (Platalea ajaja) at its nesting colony site, in early morning light. St. Johns Management Area, Florida, USA, April.

Espen Bergersen / Killer whales / orcas (Orcinus orca). Spyhopping. Kvaloya, Troms, Norway October

Espen Bergersen / Killer whales / orcas (Orcinus orca). Spyhopping. Kvaloya, Troms, Norway October

Francisco Marquez / Greater flamingo (Phoenicopterus ruber) juveniles and a few adults kept temporarily in an enclosure for ringing, Fuente de Piedra lagoon, Malaga, Spain. August.

Francisco Marquez / Greater flamingo (Phoenicopterus ruber) juveniles and a few adults kept temporarily in an enclosure for ringing, Fuente de Piedra lagoon, Malaga, Spain. August.

Francisco Marquez / Volunteers rounding up Greater flamingo (Phoenicopterus ruber) juveniles for ringing, Fuente de Piedra lagoon, Malaga, Spain. August.

Francisco Marquez / Greater flamingo (Phoenicopterus ruber) flock silhouetted in wetland, Fuente de Piedra lagoon, Malaga, Spain. August.

Jack Dykinga / Sandhill cranes (Antigone canadensis) in flight in late evening light, Bosque Del Apache, New Mexico, USA. December

Jack Dykinga / Flocks of Mallard ducks (Anas platyrhynchos) taking off in tree-lined wetlands. Bosque del Apache National Wildlife Refuge, New Mexico, USA.

Jack Dykinga / Yellow-headed blackbirds (Xanthocephalus xanthocephalus) and Redwing blackbirds (Agelaius phoeniceus) roosting. Whitewater Draw Wildlife Area, Arizona, USA, December.

Jack Dykinga / Murmuration of Yellow-headed blackbirds (Xanthocephalus xanthocephalus) and Redwing blackbirds (Agelaius phoeniceus) at sunset, before roosting. Whitewater Draw Wildlife Area, Arizona, USA, December.

SCOTLAND: The Big Picture / Red squirrel, (Sciurus vulgaris), three animals backlit on pine branch, Cairngorms National Park, Scotland, UK.May

SCOTLAND: The Big Picture / Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK., March

SCOTLAND: The Big Picture / Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April.

SCOTLAND: The Big Picture / Grey seal (Halichoerus grypus) pup on a breeding beach, Orkney, Scotland, UK, April.

SCOTLAND: The Big Picture / Grey seal (Halichoerus grypus) huge group hauled out on a beach, Island of Mingulay, west coast of Scotland. May.

SCOTLAND: The Big Picture / Grey seal (Halichoerus grypus) using freshwater pools along the shore line and in the adjacent fields as place to learn to swim, Orkney, Scotland, UK, April.

SCOTLAND: The Big Picture / Young grey seal pup (Halichoerus grypus) recently born on a beach in Orkney, Scotland, UK, April.

SCOTLAND: The Big Picture / Close up of a colony of Jewel anemones (Corynactis viridis) Scotland, UK, October.

SCOTLAND: The Big Picture / Orca (Orcinus orca) surfacing and blowing, Shetland, Scotland, UK, August.

Sandesh Kadur / Slow motion clip of Amur falcons (Falco amurensis) congregating at roost site during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Sandesh Kadur / Amur falcons (Falco amurensis) congregating during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Sandesh Kadur / Amur falcons (Falco amurensis) congregating during their migration from Siberia to Africa, near Doyang reservoir, Nagaland, India, October.

Espen Bergersen / Aerial shot of a fishing boat catching Atlantic herring (Clupea harenguis), with Killer whales (Orcinus orca) and Humpback whales (Megaptera novaeangliae) swimming around the boat and large numbers of...

Espen Bergersen / Aerial shot of a fishing boat catching Atlantic herring (Clupea harenguis), with Killer whales (Orcinus orca) and Humpback whales (Megaptera novaeangliae) swimming around the boats and large numbers o...

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming inside a shoal of Atlantic herring (Clupea harenguis), hunting, with one Killer whale tail slapping to stun the Herring, Sommaroy, Troms, Norwa...

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming into a huge shoal of Atlantic herring (Clupea harenguis), hunting, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming near to a huge shoal of Atlantic herring (Clupea harenguis), hunting, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming inside a shoal of Atlantic herring (Clupea harenguis), hunting, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) swimming inside a large shoal of Atlantic herring (Clupea harenguis), hunting, Sommaroy, Troms, Norway, January.

Espen Bergersen / Aerial shot of a huge shoal of Atlantic herring (Clupea harenguis), with a Humpback whale (Megaptera novaeangliae) nearby, Sommaroy, Troms. Norway, January.

Espen Bergersen / Aerial shot of Killer whale (Orcinus orca) hunting Atlantic herring (Clupea harengus), Troms, Norway, October.

Espen Bergersen / Aerial shot of a pod of Killer whales (Orcinus orca) swimming near the surface, Troms, Norway, October.

Espen Bergersen / Aerial shot of five Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of a group of Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, one whale creating a bubble n...

Espen Bergersen / Aerial shot of five Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus), Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of three Humpback whales (Megaptera novaeangliae) swimming into a shoal of Atlantic herring (Clupea harengus), preparing to lunge feed, Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) lunge feeding in a shoal of Atlantic herring (Clupea harengus) with some fish jumping out of the water, Troms, Norway, January.

Espen Bergersen / Aerial shot of four Humpback whales (Megaptera novaeangliae) swimming near to a shoal of Atlantic herring (Clupea harengus), Troms, Norway, January.