Ingo Arndt / Cape Gannet (Sula capensis) perched on a rock whilst calling, nesting colony, Lamberts Bay, South Africa.

Ingo Arndt / Two Northern elephant seal (Mirounga angustirostris) males fighting for dominance, California, USA

Andy Trowbridge / Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July.

Erlend Haarberg / Brambling (Fringilla montifringilla) and Chaffinch (Fringilla coelebs) fighting. Vauldalen, Sor-trondelag. Norway, May.

Michael Pitts / Gentoo penguins (Pygoscelis papua) calling and defending their eggs from Southern Skua (Stercorarius antarcticus) hunting for unattended eggs or chicks, Sea Lion island, The Falklands, South Atlantic.

Chris & Monique Fallows / Blue Shark (Prionace glauca) feeding on Anchovy (Engraulis encrasicolus) bait ball, Cape Point, South Africa.

Eric Baccega / Male Steller sea lion (Eumetopias jubatus) with a group of females hauled out at a rookery, Prince Rupert, British Columbia, Canada, June.

Ben Cranke,Ben Cranke / Gentoo Penguin (Pygoscelis papua) group travelling out to sea, Cuverville Island, Antarctic Peninsula, Antarctica

Laurent Geslin / Two male urban Red foxes, mouths open, one in submissive posture {Vulpes vulpes} London, UK

Laurent Geslin / Two dominant male urban Red foxes fighting on road, mouths open {Vulpes vulpes} London, UK

Andrew Parkinson / Gannet (Morus bassanus) chick in a line of adults. Shetland Islands, Scotland, UK, August.

Denis-Huot / Grant's zebra (Equus burchelli boehmi) in a migration herd of wildebeest (Connochaetes taurinus) Masai-Mara Game Reserve, Kenya