Ingo Arndt / Bark beetle (Scolytinae) galleries in dead spruce tree (Picea abies), Bayerischer Wald / Bavarian Forest National Park, Germany., June.

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) infant male 'Gizmo' aged 2 years, portrait. Gombe National Park, Tanzania.

Gabriel Rojo / Patagonian cavy (Dolichotis patagonum ) rear view of two, La Pampa, Argentina.

Chris & Monique Fallows / King penguin (Aptenodytes patagonicus) colony at sunrise. Grytviken, South Georgia Island.

Alex Hyde / Long exposure of light trail from bioluminescent click beetle (Pyrophorus sp) walking on ground at night. Danum Valley Sabah, Borneo.

Markus Varesvuo / Steller's Eagle (Haliaeteus pelagicus) flying, with black background, Hokkaido, Japan, February

Sergey Gorshkov / Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September.

Andy Trowbridge / Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July.

Franco Banfi / Freshwater Dorado (Salminus brasiliensis) Recanto Ecologico, Rio da Prata, Bonito, Mato Grosso do Sul, Brazil

David Pike / Harvest mouse (Micromys minutus) on stalk, West Country Wildlife Photography Centre, captive, June.

Bruno D'Amicis / Apennine chamois (Rupicapra pyrenaica ornata) male moving on extremely steep and smooth rock wall. Endemic to the Apennine mountains. Sibillini NP. Marche, Italy, August.

Eric Medard / Grey heron (Ardea cinerea) at night, taken with infra-red, Mayenne, Pays de la Loire, France, September.