Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition

Anup Shah / African fish eagle {Haliaeetus vocifer} defends Lesser flamingo carcass from Marabou storks {Leptoptilos crumeniferus}, Lake Nakuru, Kenya.

Anup Shah / African fish eagle {Haliaeetus vocifer} fighting Marabou storks {Leptoptilos crumeniferus} over Lesser flamingo carcass, Lake Nakuru, Kenya.

John Sparks / David Attenborough with mountain gorillas, on location during filming for BBC 'Life on Earth' series in Rwanda 1978

Marie Read / Western Grebe (Aechmorphus occidentalis), pair during 'rushing' courtship display in which they run across water's surface in synchrony, near Escondido, California, USA, January.

Marie Read / Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October.

Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.

Ingo Arndt / King Penguin (Aptenodytes patagonicus) dispute on beach, Volunteer Point, Falkland Islands, February

Ingo Arndt / King Penguins (Aptenodytes patagonicus) dispute while coming ashore, Salisbury Plain, South Georgia Island, February

Ingo Arndt / Thick billed ravens interacting {Corvus crassirostris} Simien NP, Ethiopia, East Africa

Mark Hamblin / 2020VISION / Two male Black grouse (Tetrao tetrix) fighting at lek site, with mountain backdrop, Cairngorms NP, Scotland, UK, April 2010

Mark Hamblin / 2020VISION / Jackdaw (Corvus monedula) pair at their nest hole, one pecking the other for attention, the other quivering its tail, Inverness-shire, Scotland, April 2011

Gavin Hellier / Ceremonially decorated Indian elephant (Elephas maximus) and mahout outside the Hawa Mahal / Palace of Winds, Rajasthan, Jaipur, India, 2011.

Ammonite / Group of African lionesses (Panthera leo) greeting each other in the rain, Masai Mara NP, Kenya

Ammonite / African lioness (Panthera leo) growling and swatting at a sub-adult male approaching a kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.

Jabruson Motion / THIS VIDEO CLIP WILL BE AVAILABLE TO VIEW ONLINE SOON. TO VIEW NOW, PLEASE CONTACT US. - Family of African forest elephants (Loxodonta africana cyclotis) gathered at a mineral dig, female reaching out...

Jabruson Motion / THIS VIDEO CLIP WILL BE AVAILABLE TO VIEW ONLINE SOON. TO VIEW NOW, PLEASE CONTACT US. - African forest elephant (Loxodonta africana cyclotis) calf bathing in and running through a waterhole, with mot...

Ammonite / Water buffalo (Bubalus arnee) chasing Sri Lankan leopard (Panthera pardus kotiya) near a waterhole, footage taken at night using thermal camera technology, Yala National Park, Sri Lanka.

Ammonite / Family of White-throated / White-faced capuchins (Cebus capucinus) resting and grooming each other, Santa Rosa National Park, Costa Rica.

Ramon Navarro / Adult Hoopoe (Upupa epops) feeding insect prey to chick at nest hole, Seville, Spain, May.

Jabruson Motion / African forest elephant (Loxodonta africana cyclotis) calf keeping larger juvenile from gaining access to a mineral dig, Zanga-Ndoki National Park, Sangha-Mbaere Prefecture, Central African Republic

Will Burrard-Lucas / Meerkat (Suricata suricatta) adult with babies, Makgadikgadi Pans, Botswana.

Laurent Geslin / Rock dove / Feral pigeon (Columba livia) pair displaying, London, UK

Laurent Geslin / Rock dove / Feral pigeon (Columba livia) pair interacting, London, UK

Laurent Geslin / Rock dove / Feral pigeon (Columba livia) pair interacting, London, UK

Laurent Geslin / Man with a feral pigeon (Columba livia) on his shoulder. Trafalgar Square, London, UK

Laurent Geslin / Black backed jackal, submission {Canis mesomelas} Skeleton Coast, Namibia

Laurent Geslin / Urban Red fox {Vulpes vulpes} + angry domestic cat in church graveyard, London, UK

Diane McAllister,Diane McAllister / Japanese Macaque (Macaca fuscata) juvenile appears to be doing tricks to get some attention in Jigokudani, Japan.

Doug Perrine / Hawaiian monk seals (Monachus schauinslandi) aggressive male attacks a five and a half week pup, mother (right) approaches to intervene; Papaloa Beach, Kalaupapa, Molokai, Hawaii, USA

Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July

Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July

Suzi Eszterhas,Suzi Eszterhas / Bornean Orangutan (Pongo pygmaeus) juvenile holding hands with carer. Orangutan Care Center, Borneo, Indonesia.

Suzi Eszterhas,Suzi Eszterhas / Bornean Orangutan (Pongo pygmaeus) baby climbing as it is trained to explore, looking down at carer. Orangutan Care Center, Borneo, Indonesia.

Denis-Huot / Bottlenose dolphin (Tursiops truncatus) playing in the waves of an oil tanker, Port Aransas, Texas, USA

Jabruson / African elephant (Loxodonta africana) two youngsters 'Mecca' and 'Lungunia' being fed mangos by their keeper, Garamba National Park, North Eastern Zaire, now the Democratic Republic of Congo. Remnants...

Roger Powell / Two Buzzards (Buteo buteo) fighting over food in snow, Dransfeld, Hannover, Germany, January

Roger Powell / Carrion Crow (Corvus corone) tweaking the wing of a Buzzard (Buteo buteo) to try to get it to move off the food, Dransfeld, Hannover, Germany, January

Franco Banfi / Free diver with Blue shark (Prionace glauca) Pico Island, Azores, Portugal, Atlantic Ocean, July. No release available.

Jane Burton / Baby Grey Squirrel (Sciurus carolinensis) 'kissing' a tabby kitten. NON DISPONIBLE POUR UNE UTILISATION DANS UN LIVRE JUSQU'EN 2025. MERCI DE NOUS CONTACTER POUR TOUT AUTRE UTILISATION AFIN DE POUVOIR...

David Tipling / Tourist photographing King Penguins (Aptenodytes patagonicus) Fortuna Bay, South Georgia, November 2006. No release available.

Paul Hobson / Arctic tern (Sterna paradisaea) two fighting in the air, Scotland, UK July

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) two individuals greeting, Tangkoko National Park, Sulawesi, Indonesia.

Mark Taylor / Beagle dog, Bruce, with ginger kitten, Tom. NON DISPONIBLE POUR UNE UTILISATION DANS UN LIVRE JUSQU'EN 2025. MERCI DE NOUS CONTACTER POUR TOUT AUTRE UTILISATION AFIN DE POUVOIR DEBLOQUER L'IMAGE

Tony Heald / African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012

Anup Shah / Western chimpanzee (Pan troglodytes verus) young male 'Jeje' aged 13 years being groomed by young male 'Peley' aged 12 years, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Western chimpanzee (Pan troglodytes verus) males 'Jeje' aged 13 years, 'Tua' aged 53 years and 'Peley' aged 12 years grooming, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Western chimpanzee (Pan troglodytes verus) female 'Fana' aged 54 years playing with her infant grandson 'Flanle' aged 3 years, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011. Sequence 4...

Anup Shah / Topi (Damaliscus lunatus jimela) two males sparring, with another approaching, Masai Mara National Reserve, Kenya. March

Bertie Gregory / 2020VISION / Juvenile male Peregrine falcon (Falco peregrinus) vocalising as its parent takes off, Bristol, England, UK, June.

Neil Aldridge / African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana

Mark Carwardine / Hands of a tourist leaning from boat to touch the mouth of a friendly Grey whale (Eschrichtius robustus) lying on the surface, San Ignacio Lagoon, Baja California, Mexico, April 2009

Ole Jorgen Liodden / Royal Penguin (Eudyptes schlegeli) fighting, Macquarie Island, Sub-Antarctic Australia. November.

Ole Jorgen Liodden / Royal Penguins (Eudyptes schlegeli) fighting, Macquarie Island, Sub-Antarctic Australia. November.

Steven Kazlowski / Polar bear (Ursus maritimus) sow and cub sliding on their backs over the pack ice, much like ice skating, 1002 coastal plain of the Arctic National Wildlife Refuge, Alaska, USA

Steven Kazlowski / Grizzly bear (Ursus arctos horribilis) young rare blonde (white) bear plays with a young dark coloured bear along the coast of Katmai National Park, Alaskan peninsula, USA

Alex Mustard / Whale shark (Rhincodon typus) silhouette of shark feeding on floating fish eggs (not visible) just below the surface in calm weather, with snorkeller taking photos nearby, Isla Mujeres, Quintana Roo,...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) female 'Mopambi' sitting with her playful infant 'Sopo' aged 18 months, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republi...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) female 'Mopambi' sitting with her playful infant 'Sopo' aged 18 months, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republi...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years playing with sub-adult female 'Mosoko' aged 8 years with juvenile male 'Tembo' aged 4 years watching in the backg...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) infant 'Sopo' aged 18 months running to his mother 'Mopambi' in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decem...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) infant 'Sopo' aged 18 months running to his mother 'Mopambi' in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decem...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) infant 'Sopo' aged 18 months running to his mother 'Mopambi' in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decem...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Tembo' aged 4 years, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central A...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Tembo' aged 4 years, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central A...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Tembo' aged 4 years, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central A...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years playing with juvenile male 'Mobangi' aged 5 years, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Centr...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Mobangi' aged 5 years, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) female 'Mopambi' sitting with her playful infant 'Sopo' aged 18 months, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republi...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Tembo' aged 4 years in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Tembo' aged 4 years in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Tembo' aged 4 years in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central...

David Pattyn / Red foxes (Vulpes vulpes) two fighting, probably over territory, The Netherlands, August

David Pattyn / Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July

Steven Kazlowski / Moose (Alces alces) newborn calves playing in spring vegetation. Tony Knowles Coastal Trail, Anchorage, south-central Alaska.

Kristel Richard / Wild meerkats (Suricata suricatta) habituated to humans standing alert on local guide, Makgadikgadi Pans, Kalahari desert, Botswana.

Kristel Richard / Wild Chincoteague (Equus caballus) two breeding stallions greeting one another, Chincoteague National Wildlife Refuge, Chincoteague Island, Virginia, USA, June

Anup Shah / Olive baboon (Papio cynocephalus anubis) female grooming another females baby, Masai Mara National Reserve, Kenya, August

Anup Shah / Lioness (Panthera leo) playing with a cub aged 9-12 months, Masai Mara National Reserve, Kenya, August

Anup Shah / Lioness (Panthera leo) playing with a cub aged 9-12 months, Masai Mara National Reserve, Kenya, August

Andrew Parkinson / Redwing (Turdus iliacus) protecting a windfall apple from another bird. Derbyshire, UK, February. (Non-ex)

Andrew Parkinson / Redwings (Turdus iliacus) squabbling over an apple in snow. Derbyshire, UK, February. British Wildlife Photographer of the Year (BWPA) competition 2012, 'Animal Behaviour' category. (Non-ex) Winner of...

Andy Rouse / European hares (Lepus europaeus) boxing, UK

Andy Rouse / European hares (Lepus europaeus) boxing, UK

Alex Mustard / 2020VISION / Young Grey seal (Halichoerus grypus) playing with snorkeller, Farne Islands, Northumberland, England, UK, July. Model released.

Roberto Rinaldi / Weedy seadragon (Phylloperyx taeniolatus) with diver watching nearby, Australia. Model released

Andy Sands / Digger Wasp (Cerceris rybyensis) Female carrying two males attempting to displace one another to mate, West sussex, England, UK, July

Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia

Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia

Peter Cairns / 2020VISION / Red squirrel (Sciurus vulgaris) approaching another as it eats a nut, Cairngorms National Park, Scotland, March 2012.

Carol Walker / Two white horses of the Camargue, interacting, greeting head to head in marshes, Camargue, Southern France

Carol Walker / White horses of the Camargue with guardian, Camargue, Southern France. No release available.

Carol Walker / Two white horses of the Camargue, head to head at water, Camargue, Southern France

Mark Hamblin / 2020VISION / Goldfinches (Carduelis carduelis) squabbling over teasel seeds in winter. Hope Farm RSPB reserve, Cambridgeshire, England, UK, February. Crop of 01390677

Danny Green / 2020VISION / Two European river otters (Lutra lutra) fighting over a fish, Isle of Mull, Inner Hebrides, Scotland, UK, December