Nick Garbutt / Aggregation / bloom / swarm of Moon Jellyfish (Aurelia aurita) in the clear waters of Loch Na Keal, off the Isle of Mull, north west Scotland, UK, June, Photographed through the surface.

DOC WHITE / Blue whale underwater close-up of head and mouth feeding with pouch inflated, Mexico

Gary Bell/ Oceanwide / Grey Nurse Shark (Carcharias taurus), Solitary Islands Marine Park, New South Wales, Australia

Sandesh Kadur / Blue wildebeest (Connochaetes taurinus) crossing a river, Serengeti National Park, Tanzania.

Theo Bosboom / Large group of Common blue damselflies (Enallagma cyathigerum) including mating couples - on stems of grass, Hondenven, Tubbergen, The Netherlands. June.

Frederique Olivier / Wide angle shot of a large flock of Blue footed boobies (Sula nebouxii) taking off from the sea and diving on a bait ball, Isla Lobos de Terra, Peru.

Frederique Olivier / Wide angle shot of a large flock of Blue footed boobies (Sula nebouxii) taking off from the sea and diving on a bait ball, Isla Lobos de Terra, Peru.

Ingo Arndt / Vogelkop bowerbird (Amblyornis inornatus) decoration material (blue rotten wood) collected in order to attract females to his bower, Arfak Mountains, Irian Jaya, New Guinea, Indonesia.

Ingo Arndt / Olive ridley sea turtles (Lepidochelys olivacea) females coming ashore at twilight during an arribada (mass nesting event) to lay eggs, Pacific Coast, Ostional, Costa Rica.

Ingo Arndt / Bed of Blue mussels (Mytilus sp.) densely packed together on rock, Lamberts Bay, South Africa

Ingo Arndt / Cape Gannet (Sula capensis) perched on a rock whilst calling, nesting colony, Lamberts Bay, South Africa.

Jim Brandenburg / American Bison (Bison bison) herd on prairie with quartz, Blue Mounds State Park, Minnesota

Brent Stephenson / Polar bear (Ursus maritimus) mother and large second year cub swim across calm water, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species.

Franco Banfi / Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica.

Claudio Contreras / Firewheel (Gaillardia pulchella) flowers at twilight, Laredo Borderlands, Texas, USA. April

Wim van den Heever / Blue wildebeest (Connochaetes taurinus) herd during migration, Masai Mara, Kenya.

Anup Shah / Blue Wildebeest (Connochaetes taurinus) herd crossing Mara River, Masai Mara, Kenya

Jordi Chias / Bryde's whale (Balaenoptera edeni) and Common dolphins (Delphinus delphis) chasing small fish offshore. Santa Maria Island, Azores, Atlantic Ocean. Third Place in the Underwater Category of the Por El...

Colin Monteath/ Hedgehog House / Kayakers paddling back to cruise ship Polar Pioneer, Lemaire Channel, Antarctic Peninsula, Antarctica

Georgette Douwma / Big-eye fish (Priacanthus hamrur). Egypt, Red Sea.

Jack Dykinga / Lightning storm with rainbow over Saguaro cactus (Carnegiea gigantea) Avra Valley, Saguaro National Park, Tucson Mountains, Arizona, USA, August 2015. Long exposure with lightning trigger.

Franco Banfi / Cory's shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal, Atlantic...

Franco Banfi / Cory's shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal, Atlantic...

Theo Allofs / Blue Wildebeest (Connochaetes taurinus) herd migrating through savanna, Masai Mara, Kenya

Claudio Contreras / Caribbean flamingo (Phoenicopterus ruber) preparing to sleep, Ria Lagartos Biosphere Reserve, Yucatan Peninsula Mexico, July. Bookplate.

Claudio Contreras / Caribbean flamingo (Phoenicopterus ruber) adults guarding chick, Ria Lagartos Biosphere Reserve, Yucatan Peninsula Mexico, June. Bookplate.

Bence Mate / Little egret (Egretta garzetta) and Grey herons (Ardea cinerea) reflected in lake at twilight, Lake Csaj, Pusztaszer, Hungary. June. Winner of the Portfolio category of the Terre Sauvage Nature Images...

Juergen Freund / Little red flying-fox (Pteropus scapulatus) flying around and roosting in trees, Atherton Tablelands, Queensland, Australia. December. Second Place in the Man and Nature portfolio category of the Terr...

Juergen Freund / Schooling silversides (Atheriniformes) split level with karst limestone cliff and West Papuan fishermen in their dugout outrigger canoe. Raja Ampat, Indonesia. Second Place in the Portfolio Award of t...

Bence Mate / White-tailed Eagle (Haliaeetus albicilla) flying, with Great egret (Ardea alba) in the foreground, Lake Csaj, Pusztaszer, Hungary. February. Winner of the Portfolio category of the Terre Sauvage Natur...

Alex Mustard / Large whip goby (Bryaninops amplus) with a parasitic copepod (with two trailing egg ribbons) perched on Red whip coral. Whale Rock. Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram S...

Mark MacEwen / Snare's island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand.

Dave Watts / Hermit ibis (Geronticus eremita) flock in flight, Morocco. Critically endangered.

Juan Carlos Munoz / Palm tree branch dipping towards water, Puerto Jimenez, Golfo Dulce, Osa Peninsula, Costa Rica.

Juan Carlos Munoz / Honeybee (Apis melifera) visiting Lavender (Lavendula angustifolia) in lavender fields, Valensole Plateau, Alpes Haute Provence, France, July.

Wim van den Heever / Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa. Runner up in the Animals in their Environment Category of the Wildl...

Jordi Chias / Atolla jellyfish (Atolla wyvillei) with Gentoo penguin (Pygoscelis papua) in the background near the surface. Antarctic Peninsula. Finalist in the Underwater category of the Wildlife Photographer of t...

Pete Oxford / Chilean flamingo (Phoenicopterus chilensis) in Lago Azul with Torres del Paine, Torres del Paine National Park, Patagonia, Chile.

David Tipling / Common frog (Rana temporaria) and spawn in pond. West Runton, North Norfolk, England, UK, March. Highly commended in the Habitat category of the BWPA Awards 2015.

Espen Bergersen / Common guillemot (Uria aalge) flock, circling outside the breeding colony at Hornoya birdcliff, Finnmark, Norway.

Claudio Contreras / Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City, Mexic...

Erlend Haarberg / Arctic fox (Alopex lagopus) cubs looking curiously, Dovrefjell-Sunndalsfjella National Park, Norway, July.

Tui De Roy / Little blue / fairy penguin (Eudyptula minor) walking on pathway to nesting burrows, Neck Game Reserve, Bruny Island, Tasmania, Australia, December.

Staffan Widstrand / American Bison (Bison bison) herd in habitat, Yellowstone National Park, Wyoming, USA, October.

Anup Shah / Burchell's Zebra (Equus burchellii) and Blue Wildebeest (Connochaetes taurinus) herd crossing river, Mara River, Masai Mara, Kenya

Frederique Olivier / Adelie penguins (Pygoscelis adeliae) wide angle portrait of two with larger group in background, Antarctica.

Frederique Olivier / Adelie penguin (Pygoscelis adeliae) group, Antarctica.

Denis-Huot / Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya.

Denis-Huot / Rear view of Wildebeest (Connochaetes taurinus) herd crossing the Mara river, Masai-Mara game reserve, Kenya.

Pascal Kobeh / Group of Short-finned pilot whales (Globicephala macrorhynchus) swimming down into deep water, Costa Rica. Pacific ocean.

Ben Cranke / Hyacinth Macaw (Anodorhynchus hyacinthinus) group grooming and feeding on palm nuts, Piaui, Brazil. July.

Sven Zacek / Roe deer (Capreolus capreolus) in fog at sunset, Estonia, September. Nominated in the Melvita Nature Images Awards competition 2014.

Hougaard Malan / Large wave crashing against rock shelf. Luphathana, Pondoland, Eastern Cape, South Africa. June 2012.

Markus Varesvuo / Guillemot (Uria aalge) flock in flight over nesting colony on rocks, Norway, July 2008

Alex Mustard / Large female Southern stingray (Hypanus americanus) in shallow water, under storm clouds, Stingray City, Grand Cayman. Stingray City is feeding site for stingrays and bait was used on this shoot. Caym...

Bence Mate / Grey herons (Ardea cinerea) standing in water a dawn, Puszaszer, Hungary, July.

David Pattyn / Flowering Thrift (Armeria maritima) photographed on Great Saltee with a view of the island behind. Great Saltee, Saltee Islands, Co. Wexford, Republic of Ireland, June.

Inaki Relanzon / Zebu cattle (Bos indicus) market in Ambalavao, central Madagascar, November 2012.

Anup Shah / Eastern white-bearded wildebeest (Connochaetes taurinus) herd crossing the Mara River watched by tourists. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Common or plains zebra (Equus quagga burchelli) and Eastern white-bearded wildebeest (Connochaetes taurinus) mixed herd running. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera...

Anup Shah / Eastern white-bearded wildebeest (Connochaetes taurinus) herd running. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Jussi Murtosaari / Jackdaw (Corvus monedula) flockin flight with two Hooded Crows (Corvus cornix) and the moon, central Finland, March.

Chris & Monique Fallows / Indo Pacific Sailfish (Istiophorus platypterus) feeding on sardines (Sardina pilchardus), Isla Mujeres, Mexico, March.

Michele Westmorland / Large school of scalloped hammerhead sharks (Sphyrna lewini). Galapagos Islands.

Mark Taylor / Liver Flatcoated Retriever puppy, 6 weeks.

Tom Gilks / Fishermen preparing nets, Remba Island, Lake Victoria, Kenya.

Chris & Monique Fallows / Cape Gannet (Morus capensis) feeding on sardines, Port St Johns, Transkei, South Africa.

Chris & Monique Fallows / Indo Pacific Sailfish (Istiophorus platypterus) feeding on sardines, Isla Mujeres, Mexico.

Chris & Monique Fallows / Indo Pacific Sailfish (Istiophorus platypterus) feeding on sardines, Isla Mujeres, Mexico.

Chris & Monique Fallows / Blue Shark (Prionace glauca) feeding on Anchovy (Engraulis encrasicolus) bait ball, Cape Point, South Africa.

Denis-Huot,Denis-Huot / Lesser flamingo (Phoeniconaias minor) aerial view of flock and geyser, Bogoria Game Reserve, Kenya

Alex Mustard,Alex Mustard / A blue light fluorescence photo of Jewel anemones (Corynactis viridis). The blue light source excites the fluorescent pigments in the anemones, which emit light of other colours. The muddy seabed and...

Franco Banfi / Diver with rebreather inside sea cage used for aquaculture with thousand of Gilt-head bream (Sparus aurata) Ponza Island, Italy, Tyrrhenian Sea, Mediterranean. Model released.

Franco Banfi / Diver with rebreather inside sea cage used for aquaculture with thousand of Gilt-head bream (Sparus aurata) Ponza Island, Italy, Tyrrhenian Sea, Mediterranean. Model released.

Franco Banfi / Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean

Theo Bosboom,Theo Bosboom / Gulls, predominantly Iceland gulls (Larus glaucoides) flying above stormy ocean near Vik, Iceland, March 2012.

Theo Bosboom,Theo Bosboom / Common blue damselflies (Enallagma cyathigerum) flying over the water to rest on plant stem, in a fen, Hondeven, Holland, July

MIKE POTTS,Mike Potts / Stellers Eider (Polysticta stelleri) a small group in flight, near Vardo. North Norway, March.

Eric Dragesco,Eric Dragesco / Herd of Tibetan Wild Asses / Kiang (Equus kiang) ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India, October 2012

Andrew Parkinson / Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September.

Yukihiro Fukuda / Japanese fireflies (Luciola cruciata) in flight at night, Japan endemic species, Ichikawa-Riveer, Asago Hyougo, Japan, July

Yukihiro Fukuda,Yukihiro Fukuda / Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May

Juan Carlos Munoz / Double-crested Cormorants (Phalacrocorax auritus) and Brown pelicans (Pelecanus occidentalis) on small island, Isla Contoy National Park, Quintana Roo, Yuccatan Penninsula, Mexico

Anup Shah,Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd on the move Masai Mara National Reserve, Kenya, July

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River and clambering up steep bank. Masai Mara National Reserve, Kenya, July

Denis-Huot,Denis-Huot / Grant's zebra (Equus burchelli boehmi) herd with large cloud above, Amboseli National Park, Kenya

Denis-Huot,Denis-Huot / Grant's zebra (Equus burchelli boehmi) in a migration herd of wildebeest (Connochaetes taurinus) Masai-Mara Game Reserve, Kenya

Denis-Huot / Grant's zebra (Equus burchelli boehmi) in a migration herd of wildebeest (Connochaetes taurinus) Masai-Mara Game Reserve, Kenya

Juan Carlos Munoz,Juan Carlos Munoz / Lesser flamingo (Phoeniconaias minor) flock in lake in morning, Lake Nakuru NP, Rift Valley, Kenya

Michael Pitts,Michael Pitts / Sooty Terns (Onychoprion fuscata) flock in flight, Midway Island. Central Pacific.

Michael Pitts,Michael Pitts / Coral wall with Anthiinae schooling fish, Red Sea

Sue Daly,Sue Daly / Blue Maomao (Scorpis violaceus) shoal, Poor Knights Islands, New Zealand, February

Sue Daly / Koheru (Decapterus koheru) shoal swiming around diver, Cavalli Islands, New Zealand, February 2013. Model released.

Barrie Britton / King Penguin (Aptenodytes patagonicus) colony, with mountains reflected in the ocean, St Andrew's Bay, South Georgia. Photograph taken on location for the BBC Frozen Planet series, October 2009.

Mark Carwardine / Aerial view of Blue whale + calf {Balaenoptera musculus} Sea of Cortez, Mexico

Mark Carwardine / Aerial view of blue whale {Balaenoptera musculus}, Sea of Cortez, Mexico