John Abbott / Glorious scarab (Chrysina gloriosa) flying, Brewster County, Texas, USA. Controlled conditions. August

Robert Thompson / Oleander Hawkmoth (Daphnis nerii) resting, Southern Sicily, Italy. March.

Nick Garbutt / Malachite Butterfly (Siproeta stelenes) hanging upside down on leaf, forest near Napo River, Amazonia, Ecuador, South America.

Ross Hoddinott / Male banded demoiselle (Calopteryx splendens), resting on reed, Lower Tamar Lakes, Cornwall, UK. June.

Jussi Murtosaari / Speckled Wood (Pararge aegeria) male in habitat, Finland, April.

Jack Dykinga / Scarlet Macaw (Ara macao) in tropical rainforest canopy with beech almond fruit. Osa Peninsula, Costa Rica

Paul Harcourt Davies / Newly emerged adult Two-tailed Pasha (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August

Paul Harcourt Davies / Newly emerged adult Two-tailed Pasha butterfly (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August

Paul Harcourt Davies,Paul Harcourt Davies / Emergence of Two-tailed Pasha butterfly (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August. Sequence 6 of 9

Paul Harcourt Davies / Emergence of Two-tailed Pasha butterfly (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August. Sequence 1 of 9

Benjamin Barthelemy / Large white butterfly (Pieris brassicae), Vallee de l'Eure (Eure Valley), Eure-et-Loir, France, August

Guy Edwardes / Superb Starling (Lamprotornis superbus) with wing raised as a threat to nearby rival. Serengeti National Park, Tanzania, Africa, February