Bence Mate / Leaf cutter ant (Atta sp) female worker carrying leaf to nest, with a smaller 'minor' ant riding. The minor's job is to protect her sister from parasitic flies. Costa Rica, December. Highly Commended,...

Tui De Roy / Blue-footed Booby (Sula nebouxii) performing foot-lifting courtship display, Seymour Island, Galapagos Islands, Ecuador

Tui De Roy / Blue-footed Booby (Sula nebouxii) performing foot-lifting courtship display, Seymour Island, Galapagos Islands, Ecuador

Bence Mate / Great egret (Ardea alba) hunting fish, Lake Csaj, Pusztaszer, Hungary, February. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015.

Tony Heald / Asian elephant (Elephas maximus) domesticated elephant working, picking reeds, Kaziranga National Park, Assam, India, March 2009.

Cyril Ruoso / African elephant (Loxodonta africana) hanging upside down, during translocation by Kenya Wildlife Service, due to over population, Mwaluganje Reserve, Kenya. October 2006.

Eric Baccega / Men from the Mursi tribe decoratively scaring skin, one man scars the arm of another one by lifting the skin with an acacia spine and cutting it with a razor blade, Ethiopia, March 2015.

Espen Bergersen / Gyrfalcon (Falco rusticolus) flying with guillemot prey. Hornoya bird cliff. Finnmark, Norway. March

Chris & Monique Fallows / Orca male (Orcinus orca) raising pectoral flipper out of the water, Prince William Sound, Alaska, USA.

Chris Mattison,Chris Mattison / Rubber Boa (Charina bottae) in defensive posture, raising tail to mimic head. California, USA, November.

Anup Shah / Spotted hyena (Crocuta crocuta) preparing to carry a pup aged 2-3 months. Masai Mara National Reserve, Kenya, July

Shibai Xiao / State-owned forestry plant with workers lifting logs early in the morning, Heilongjiang Provinc, China, January 2010. No release available