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Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using 'Starlight Camera' technology without artificial lighting.
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Martin Dohrn / Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using 'Starlight Camera' technology without artificial lighting.
Lesser Flamingo (Phoenicopterus minor) wading, Amboseli National Park, Kenya
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Chase Dekker / Lesser Flamingo (Phoenicopterus minor) wading, Amboseli National Park, Kenya
Lesser Flamingo (Phoenicopterus minor) pair fighting, Amboseli National Park, Kenya
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Chase Dekker / Lesser Flamingo (Phoenicopterus minor) pair fighting, Amboseli National Park, Kenya
African Wild Dog (Lycaon pictus) group attacking Cape Warthog (Phacochoerus aethiopicus), Tswalu Game Reserve, South Africa
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Juergen & Christine Sohns / African Wild Dog (Lycaon pictus) group attacking Cape Warthog (Phacochoerus aethiopicus), Tswalu Game Reserve, South Africa
Australian Giant Cuttlefish (Sepia apama) group gathered for mating, Whyalla, South Australia, Australia
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Gary Bell/ Oceanwide / Australian Giant Cuttlefish (Sepia apama) group gathered for mating, Whyalla, South Australia, Australia
Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December
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Oscar Dewhurst / Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December
African Elephant (Loxodonta africana) herd drinking at waterhole, Mashatu Game Reserve, Botswana
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Suzi Eszterhas / African Elephant (Loxodonta africana) herd drinking at waterhole, Mashatu Game Reserve, Botswana
African Elephant (Loxodonta africana) herd drinking at waterhole, Mashatu Game Reserve, Botswana
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Suzi Eszterhas / African Elephant (Loxodonta africana) herd drinking at waterhole, Mashatu Game Reserve, Botswana
Przewalski's Horse (Equus ferus przewalskii) herd, Gobi Desert, Mongolia
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Cyril Ruoso / Przewalski's Horse (Equus ferus przewalskii) herd, Gobi Desert, Mongolia
Przewalski's Horse (Equus ferus przewalskii) herd, Gobi Desert, Mongolia
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Cyril Ruoso / Przewalski's Horse (Equus ferus przewalskii) herd, Gobi Desert, Mongolia
African Elephant (Loxodonta africana) herd on savanna, Mount Kilimanjaro, Amboseli National Park, Kenya
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Jami Tarris / African Elephant (Loxodonta africana) herd on savanna, Mount Kilimanjaro, Amboseli National Park, Kenya
Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018.
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Daniel Heuclin / Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018.
Northern Pintail (Anas acuta) males flying with female in spring courtship display, Montana
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Donald M. Jones / Northern Pintail (Anas acuta) males flying with female in spring courtship display, Montana
Humpback Whale (Megaptera novaeangliae) pod cooperative bubble-net feeding, Frederick Sound, Alaska
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Paul Souders/ Worldfoto / Humpback Whale (Megaptera novaeangliae) pod cooperative bubble-net feeding, Frederick Sound, Alaska
Humpback Whale (Megaptera novaeangliae) pod cooperative bubble-net feeding, Frederick Sound, Alaska
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Paul Souders/ Worldfoto / Humpback Whale (Megaptera novaeangliae) pod cooperative bubble-net feeding, Frederick Sound, Alaska
Long-beaked Common Dolphin (Delphinus capensis) pod porpoising, Monterey Bay, California
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Chase Dekker / Long-beaked Common Dolphin (Delphinus capensis) pod porpoising, Monterey Bay, California
African Elephant (Loxodonta africana) smelling, Masai Mara, Kenya
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Anup Shah / African Elephant (Loxodonta africana) smelling, Masai Mara, Kenya
African Elephant (Loxodonta africana) herd, Amboseli National Park, Kenya
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Hiroya Minakuchi / African Elephant (Loxodonta africana) herd, Amboseli National Park, Kenya
Asian Elephant (Elephas maximus) juvenile crossing road at night in headlight of car watched by people on the road, West Bengal, India
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Avijan Saha/ BIA / Asian Elephant (Elephas maximus) juvenile crossing road at night in headlight of car watched by people on the road, West Bengal, India
Gray Wolf (Canis lupus) pack on ridge in winter, Lamar Valley, Yellowstone National Park, Wyoming
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Daniel J. Cox / Gray Wolf (Canis lupus) pack on ridge in winter, Lamar Valley, Yellowstone National Park, Wyoming
Long-tailed Macaque (Macaca fascicularis) troop, Khao Sam Roi Yot National Park, Thailand
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Cyril Ruoso / Long-tailed Macaque (Macaca fascicularis) troop, Khao Sam Roi Yot National Park, Thailand
Spike-browed Frog (Sphenophryne cornuta) father transporting froglets, Digul River, West Papua, Indonesia
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Chien Lee / Spike-browed Frog (Sphenophryne cornuta) father transporting froglets, Digul River, West Papua, Indonesia
Spike-browed Frog (Sphenophryne cornuta) father transporting froglets, Digul River, West Papua, Indonesia
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Chien Lee / Spike-browed Frog (Sphenophryne cornuta) father transporting froglets, Digul River, West Papua, Indonesia
Dromedary (Camelus dromedarius) caravan transporting salt across salt flat, Lake Asale, Danakil Depression, Ethiopia
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Vincent Grafhorst / Dromedary (Camelus dromedarius) caravan transporting salt across salt flat, Lake Asale, Danakil Depression, Ethiopia
Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in mud at night, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
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Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in mud at night, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in mud at night, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
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Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in mud at night, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in seasonal pond, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
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Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in seasonal pond, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in seasonal pond, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
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Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in seasonal pond, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
African Elephant (Loxodonta africana) herd in plain, Masai Mara, Kenya
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Anup Shah / African Elephant (Loxodonta africana) herd in plain, Masai Mara, Kenya
King Penguin (Aptenodytes patagonicus) group on beach, Volunteer Beach, East Falkland Island, Falkland Islands
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Tui De Roy / King Penguin (Aptenodytes patagonicus) group on beach, Volunteer Beach, East Falkland Island, Falkland Islands
Nazca Booby (Sula granti) group on coast, Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
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Tui De Roy / Nazca Booby (Sula granti) group on coast, Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Nazca Booby (Sula granti) group on coast, Wolf Island, Galapagos Islands, Ecuador
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Tui De Roy / Nazca Booby (Sula granti) group on coast, Wolf Island, Galapagos Islands, Ecuador
Blue-footed Booby (Sula nebouxii) males courting female, Punta Vicente Roca, Isabela Island, Galapagos Islands, Ecuador
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Tui De Roy / Blue-footed Booby (Sula nebouxii) males courting female, Punta Vicente Roca, Isabela Island, Galapagos Islands, Ecuador
Leafcutter Ant (Atta sp) group harvesting leaf, Yasuni National Park, Ecuador
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Paul Bertner / Leafcutter Ant (Atta sp) group harvesting leaf, Yasuni National Park, Ecuador
Group of Common frogs (Rana temporaria) spawning in a garden pond, with floating spawn, Somerset, England, UK, March.
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John Waters / Group of Common frogs (Rana temporaria) spawning in a garden pond, with floating spawn, Somerset, England, UK, March.
Greater Flamingo (Phoenicopterus ruber) flock flying, Los Flamencos Sanctuary, Guajira Peninsula, Colombia
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Thomas Marent / Greater Flamingo (Phoenicopterus ruber) flock flying, Los Flamencos Sanctuary, Guajira Peninsula, Colombia
Guira Cuckoo (Guira guira) group huddled together, native to South America
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Rosl Roessner/ BIA / Guira Cuckoo (Guira guira) group huddled together, native to South America
Asian Elephant (Elephas maximus) herd crossing road, West Bengal, India
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Avijan Saha/ BIA / Asian Elephant (Elephas maximus) herd crossing road, West Bengal, India
White-fronted Goose (Anser albifrons) group flying, North Rhine-Westphalia, Germany
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Hans Glader/ BIA / White-fronted Goose (Anser albifrons) group flying, North Rhine-Westphalia, Germany
Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December.
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Yashpal Rathore / Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December.
African Elephant (Loxodonta africana) herd during storm, Masai Mara, Kenya
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Anup Shah / African Elephant (Loxodonta africana) herd during storm, Masai Mara, Kenya
African Elephant (Loxodonta africana) herd during storm, Masai Mara, Kenya
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Anup Shah / African Elephant (Loxodonta africana) herd during storm, Masai Mara, Kenya
False killer whale (Pseudorca crassidens) pod, off the Kona Coast of Hawaii Island, Hawaiian Islands, USA. January.
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Doug Perrine / False killer whale (Pseudorca crassidens) pod, off the Kona Coast of Hawaii Island, Hawaiian Islands, USA. January.
Great egret (Ardea alba) group of five at water's edge, with very out of focus egret in foreground. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, February.
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Bence Mate / Great egret (Ardea alba) group of five at water's edge, with very out of focus egret in foreground. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, February.
Rockhopper Penguin (Eudyptes chrysocome) group coming ashore, Falkland Islands
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Heike Odermatt / Rockhopper Penguin (Eudyptes chrysocome) group coming ashore, Falkland Islands
Group of Sperm whales (Physter macrocephalus) family group swimming together, Indian Ocean, March.
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Tony Wu / Group of Sperm whales (Physter macrocephalus) family group swimming together, Indian Ocean, March.
Humpback Whale (Megaptera novaeangliae) group gulp feeding, Alaska
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Matthias Breiter / Humpback Whale (Megaptera novaeangliae) group gulp feeding, Alaska
Trumpeter Swan (Cygnus buccinator) flock flying, Arkansas
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Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) flock flying, Arkansas
Cheetah (Acinonyx jubatus) male walking by Burchell's Zebra (Equus burchellii) herd, Masai Mara, Kenya
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Ingo Arndt / Cheetah (Acinonyx jubatus) male walking by Burchell's Zebra (Equus burchellii) herd, Masai Mara, Kenya
Blue wildebeest (Connochaetes taurinus) herd silhouetted against the rising sun with clouds in the background. Greater Kruger National Park, South Africa
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Wim van den Heever / Blue wildebeest (Connochaetes taurinus) herd silhouetted against the rising sun with clouds in the background. Greater Kruger National Park, South Africa
White horses of the Camargue galloping through marshes in the Camargue, France. April.
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Jeff Vanuga / White horses of the Camargue galloping through marshes in the Camargue, France. April.
Snow Goose (Chen caerulescens) group flying, central Montana
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Donald M. Jones / Snow Goose (Chen caerulescens) group flying, central Montana
Elk (Cervus elaphus) bull and harem of females and calves along river, Athabasca River, western Canada
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Donald M. Jones / Elk (Cervus elaphus) bull and harem of females and calves along river, Athabasca River, western Canada
African Elephant (Loxodonta africana) mother and calf, Amboseli National Park, Kenya
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Hiroya Minakuchi / African Elephant (Loxodonta africana) mother and calf, Amboseli National Park, Kenya
African Elephant (Loxodonta africana) herd walking, Amboseli National Park, Kenya
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Hiroya Minakuchi / African Elephant (Loxodonta africana) herd walking, Amboseli National Park, Kenya
African Elephant (Loxodonta africana) herd, Masai Mara, Kenya
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Anup Shah / African Elephant (Loxodonta africana) herd, Masai Mara, Kenya
Bryde's whale (Balaenoptera edeni) and Common dolphins (Delphinus delphis) chasing small fish offshore. Santa Maria Island, Azores, Atlantic Ocean. Third Place in the Underwater Category of the Por El...
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Jordi Chias / Bryde's whale (Balaenoptera edeni) and Common dolphins (Delphinus delphis) chasing small fish offshore. Santa Maria Island, Azores, Atlantic Ocean. Third Place in the Underwater Category of the Por El...
Harbor Seal (Phoca vitulina) group on ice floes, Prince William Sound, Alaska
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Suzi Eszterhas / Harbor Seal (Phoca vitulina) group on ice floes, Prince William Sound, Alaska
Inuit sled dogs resting, near Ittoqortoormiit, Scoresbysund, Greenland. Second Prize in the Nature category of the Spider Black and White Photography Competition 2015.
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Pal Hermansen / Inuit sled dogs resting, near Ittoqortoormiit, Scoresbysund, Greenland. Second Prize in the Nature category of the Spider Black and White Photography Competition 2015.
Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park.  South Africa. Specially commended in the Animal Portraits category of...
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Staffan Widstrand / Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa. Specially commended in the Animal Portraits category of...
Great egrets (Ardea alba) and Grey heron (Ardea cinerea) squabbling over fish, with some in flight. Blurred motion photograph, Lake Csaj, Pusztaszer, Hungary. February.
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Bence Mate / Great egrets (Ardea alba) and Grey heron (Ardea cinerea) squabbling over fish, with some in flight. Blurred motion photograph, Lake Csaj, Pusztaszer, Hungary. February.
Gelada Baboon (Theropithecus gelada) male and females huddling for warmth, Simien Mountains National Park, Ethiopia
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Thomas Marent / Gelada Baboon (Theropithecus gelada) male and females huddling for warmth, Simien Mountains National Park, Ethiopia
Group of predators hunting together, in an example of 'Nuclear foraging'. The Moray eel (Gymnothorax javanicus) flushes out prey, and is followed by a Napolean wrasse (Cheilinus undulatus) and a Bluef...
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Alex Mustard / Group of predators hunting together, in an example of 'Nuclear foraging'. The Moray eel (Gymnothorax javanicus) flushes out prey, and is followed by a Napolean wrasse (Cheilinus undulatus) and a Bluef...
Corsican pig (Sus scrofa domestica) sow, suckling the piglets, Parc Naturel Regional de Corse / Natural Regional Park of Corsica, Corsica Island, France. September.
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Pascal Pittorino / Corsican pig (Sus scrofa domestica) sow, suckling the piglets, Parc Naturel Regional de Corse / Natural Regional Park of Corsica, Corsica Island, France. September.
Ringed-tailed lemur (Lemur catta) foraging in sisal field looking for flowers during a poor year without much fruit. Berenty reserve, Madagascar, September.
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Cyril Ruoso / Ringed-tailed lemur (Lemur catta) foraging in sisal field looking for flowers during a poor year without much fruit. Berenty reserve, Madagascar, September.
Keeper feeding Common woolly monkeys (Lagothrix lagotricha) in Ikamaperou Sanctuary, Amazon, Peru. October 2006.
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Cyril Ruoso / Keeper feeding Common woolly monkeys (Lagothrix lagotricha) in Ikamaperou Sanctuary, Amazon, Peru. October 2006.
Vicuna (Vicugna vicugna) herd  in Altiplano, Aguada Blanca National Reserve, Peru.
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Cyril Ruoso / Vicuna (Vicugna vicugna) herd in Altiplano, Aguada Blanca National Reserve, Peru.
Kittiwakes (Rissa tridactyla) nesting on wall of house, Vardo town, Varanger Peninsula, Norway, March.
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Staffan Widstrand / Kittiwakes (Rissa tridactyla) nesting on wall of house, Vardo town, Varanger Peninsula, Norway, March.
Four Roe deer (Capreolus capreolus) bucks with velvet antlers and the shadow of a fifth buck, Tartumaa, Estonia, March.
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Sven Zacek / Four Roe deer (Capreolus capreolus) bucks with velvet antlers and the shadow of a fifth buck, Tartumaa, Estonia, March.
American buffalo (Bison bison) with group of calves, Yellowstone National Park, Wyoming, USA. May.
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George Sanker / American buffalo (Bison bison) with group of calves, Yellowstone National Park, Wyoming, USA. May.
Magnificent frigate bird (Fregata magnificens) group flying against cloudy sky, Galapagos.
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Andy Rouse / Magnificent frigate bird (Fregata magnificens) group flying against cloudy sky, Galapagos.
Small flock of Brent Geese (Branta bernicla) in flight..Texel Island, The Netherlands, Europe
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Roger Powell / Small flock of Brent Geese (Branta bernicla) in flight..Texel Island, The Netherlands, Europe
Crested myna starlings (Arcidotheres cristatellus) sitting on domestic waterbuffaloes (Bubalus domesticus), Poyang Ho Lake, Jiangxi province, China
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Staffan Widstrand / Wild Wonders of China / Crested myna starlings (Arcidotheres cristatellus) sitting on domestic waterbuffaloes (Bubalus domesticus), Poyang Ho Lake, Jiangxi province, China
Long-beaked common dolphin (Delphinus capensis) school near the water?s surface, False Bay, South Africa
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Chris & Monique Fallows / Long-beaked common dolphin (Delphinus capensis) school near the water?s surface, False Bay, South Africa
Red Stink Bug (Pycanum rubeus) hatchlings cluster to amplify the effect of their aposematic warning coloration, Malaysia
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Chien Lee / Red Stink Bug (Pycanum rubeus) hatchlings cluster to amplify the effect of their aposematic warning coloration, Malaysia
Kittiwakes (Rissa tridactyla) squabbling on ice. Svalbard, Norway.  July
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Espen Bergersen / Kittiwakes (Rissa tridactyla) squabbling on ice. Svalbard, Norway. July
African elephant (Loxodonta africana) herd eating, Masai-Mara Game Reserve, Kenya
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Denis-Huot / African elephant (Loxodonta africana) herd eating, Masai-Mara Game Reserve, Kenya
Galapagos penguin (Spheniscus mendiculus) group underwater, Galapagos. Endemic.
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Ole Jorgen Liodden / Galapagos penguin (Spheniscus mendiculus) group underwater, Galapagos. Endemic.
Galapagos Sea Lion (Zalophus wollebaeki) group resting on beach, Gardner Bay, Hood Island, Galapagos Islands, Ecuador
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Pete Oxford / Galapagos Sea Lion (Zalophus wollebaeki) group resting on beach, Gardner Bay, Hood Island, Galapagos Islands, Ecuador
Galapagos Penguin (Spheniscus mendiculus) group swimming, Galapagos Islands, Ecuador
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Pete Oxford / Galapagos Penguin (Spheniscus mendiculus) group swimming, Galapagos Islands, Ecuador
Arctic fox (Alopex lagopus) cubs looking curiously, Dovrefjell-Sunndalsfjella National Park, Norway, July.
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Erlend Haarberg / Arctic fox (Alopex lagopus) cubs looking curiously, Dovrefjell-Sunndalsfjella National Park, Norway, July.
Northern Flicker (Colaptes auratus) entering nest cavity with chicks, Alaska
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Michael Quinton / Northern Flicker (Colaptes auratus) entering nest cavity with chicks, Alaska
Northern Flicker (Colaptes auratus) entering nest cavity with begging chicks, Alaska
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Michael Quinton / Northern Flicker (Colaptes auratus) entering nest cavity with begging chicks, Alaska
Northern Flicker (Colaptes auratus) parent brooding chicks in nest cavity, Alaska
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Michael Quinton / Northern Flicker (Colaptes auratus) parent brooding chicks in nest cavity, Alaska
Northern Flicker (Colaptes auratus) eggs in nest cavity, Alaska
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Michael Quinton / Northern Flicker (Colaptes auratus) eggs in nest cavity, Alaska
Impala (Aepyceros melampus) herd running, Sabi-sands Game Reserve, South Africa
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Sergey Gorshkov / Impala (Aepyceros melampus) herd running, Sabi-sands Game Reserve, South Africa
Snow Goose (Chen caerulescens) group flying, central New Mexico
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Donald M. Jones / Snow Goose (Chen caerulescens) group flying, central New Mexico
Emperor Penguin (Aptenodytes forsteri) chicks, Queen Maud Land, Antarctica
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Stefan Christmann / Emperor Penguin (Aptenodytes forsteri) chicks, Queen Maud Land, Antarctica
Band of wild Mongolian horses running on plains at the foot of Dungurukh Uul mountain, near the border with China and Kazakhstan, Bayan-Olgiy aymag, Mongolia. September.
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Kristel Richard / Band of wild Mongolian horses running on plains at the foot of Dungurukh Uul mountain, near the border with China and Kazakhstan, Bayan-Olgiy aymag, Mongolia. September.
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea, close to Little Brother Island. The Brothers Islands, Egy...
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Alex Mustard / Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea, close to Little Brother Island. The Brothers Islands, Egy...
Black kite (Milvus migrans) flying and red-crowned crane (Grus japonensis) on snow, Hokkaido, Japan, February.
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Markus Varesvuo / Black kite (Milvus migrans) flying and red-crowned crane (Grus japonensis) on snow, Hokkaido, Japan, February.
Goldeneye (Bucephala clangula) with chicks swimming behind, Helsinki, Uusimaa, Finland, May.
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Jussi Murtosaari / Goldeneye (Bucephala clangula) with chicks swimming behind, Helsinki, Uusimaa, Finland, May.
Chinstrap penguin (Pygoscelis antarcticus) group on blue iceberg, Antarctica
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Andy Rouse / Chinstrap penguin (Pygoscelis antarcticus) group on blue iceberg, Antarctica
African elephant (Loxodonta africana) herd at sunset, Masai-Mara Game Reserve, Kenya. January.
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Denis-Huot / African elephant (Loxodonta africana) herd at sunset, Masai-Mara Game Reserve, Kenya. January.
African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October.
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Denis-Huot / African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October.
Orca (Orcinus orca) pod surfacing, Prince William Sound, Alaska
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Hiroya Minakuchi / Orca (Orcinus orca) pod surfacing, Prince William Sound, Alaska