Fabio Pupin / Large psammodromus, (Psammodromus algirus), male with mating season colours near stream, Spain, June . Non-ex.
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Tony Wu / Asian sheepshead wrasse (Semicossyphus reticulatus) mature male in breeding colouration. Niigata, Japan. non-ex. Awarded in Animal Portraits category, Wildlife Photographer of the Year 2018.
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Robin Chittenden / Guillemot (Uria aalge) in winter plumage, flapping its wings whilst standing in sea foam, Blakeney Point, Norfolk, UK. North Sea. November.
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Roger Powell / Grey plover (Pluvialis squatarola) standing on tidal mud. Lindisfarne Nature Reserve, Northumberland, UK. January.
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Roger Powell / Sanderling (Calidris alba) walking across coastal sand. Boulmer Beach, Northumberland, UK. January.
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Roger Powell / Two Red knots (Calidris canutus), in winter plumage, searching for food in tidal pool. Lindisfarne National Nature Reserve, Northumberland, UK. January.
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Roger Powell / Red knot (Calidris canutus), in winter plumage, consuming food from tidal pool. Lindisfarne National Nature Reserve, Northumberland, UK. January.
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Roger Powell / Red knot (Calidris canutus), in winter plumage, searching for food in tidal pool. Lindisfarne National Nature Reserve, Northumberland, UK. January.
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Tui De Roy / Swallow-tailed gull (Creagrus furcatus), in non-breeding plumage, rarely seen ashore, walking across beach, Genovesa Island, Galapagos Islands, Ecuador.
