John Cancalosi / Sonoran desert toad (Incilius alvarius) pair mating, with trail of spawn in water. Arizona, USA.

George Sanker / Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.

George Sanker / Great egret (Ardea alba) diving head into water to catch prey, in evening light. Myakka River State Park, Florida, USA, March. Sequence 1/2.

Jack Dykinga / Morning light with autumn coloured Cottonwoods (Populus sp) trees lining the river. Valley of the Gods, Colorado Plateau, of the Great Basin Desert, Utah, USA, November 2018.

Nick Hawkins / Underneath view of a North American beaver (Castor canadensis) swimming under the ice of a frozen river. Quebec, Canada. October.

SCOTLAND: The Big Picture / Buacaille Etive Mor and River Etive in winter, Rannoch Moor, Glencoe, Scotland, UK.December

Marie Read / Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her and crossing his bill with hers...

Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.

George Sanker / North American river otter (Lontra canadensis) swimming with head above water. Acadia National Park, Maine, USA. November.

Nick Garbutt / Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January

Paul Williams / Moose (Alces alces) female with twin calves, Baxter State Park, Maine, USA, June.

Jack Dykinga / Toroweap Overlook in sunrise light illuminating rim rocks with Colorado River far below, Grand Canyon National Park, Arizona, USA.